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<journal-id journal-id-type="publisher-id">Front. Sustain. Resour. Manag.</journal-id>
<journal-title-group>
<journal-title>Frontiers in Sustainable Resource Management</journal-title>
<abbrev-journal-title abbrev-type="pubmed">Front. Sustain. Resour. Manag.</abbrev-journal-title>
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<issn pub-type="epub">2813-3005</issn>
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<publisher-name>Frontiers Media S.A.</publisher-name>
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<article-id pub-id-type="doi">10.3389/fsrma.2026.1732587</article-id>
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<subj-group subj-group-type="heading">
<subject>Original Research</subject>
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<title-group>
<article-title>Institutional analysis of agroforestry in the Western Highlands of Cameroon: processes and outcomes</article-title>
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<name><surname>Nihngwo</surname> <given-names>Alei Christa</given-names></name>
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<name><surname>Awazi</surname> <given-names>Nyong Princely</given-names></name>
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<name><surname>Kimengsi</surname> <given-names>Jude Ndzifon</given-names></name>
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<aff id="aff1"><label>1</label><institution>Forest Institutions and International Development (FIID) Research Group, Chair of Tropical and International Forestry, Faculty of Environmental Sciences, Dresden University of Technology</institution>, <city>Tharandt</city>, <country country="de">Germany</country></aff>
<aff id="aff2"><label>2</label><institution>Department of Forestry and Wildlife Technology, College of Technology (COLTECH), University of Bamenda</institution>, <city>Bambili</city>, <country country="cm">Cameroon</country></aff>
<aff id="aff3"><label>3</label><institution>Department of Geography, University of Bamenda</institution>, <city>Bambili</city>, <country country="cm">Cameroon</country></aff>
<author-notes>
<corresp id="c001"><label>&#x0002A;</label>Correspondence: Nyong Princely Awazi, <email xlink:href="mailto:nyongprincely@gmail.com">nyongprincely@gmail.com</email>; <email xlink:href="mailto:awazinyong@uniba.cm">awazinyong@uniba.cm</email></corresp>
<fn fn-type="other" id="fn001"><label>&#x02020;</label><p>ORCID: Nyong Princely Awazi <uri xlink:href="https://orcid.org/0000-0002-0801-0719">orcid.org/0000-0002-0801-0719</uri></p></fn></author-notes>
<pub-date publication-format="electronic" date-type="pub" iso-8601-date="2026-02-19">
<day>19</day>
<month>02</month>
<year>2026</year>
</pub-date>
<pub-date publication-format="electronic" date-type="collection">
<year>2026</year>
</pub-date>
<volume>5</volume>
<elocation-id>1732587</elocation-id>
<history>
<date date-type="received">
<day>26</day>
<month>10</month>
<year>2025</year>
</date>
<date date-type="rev-recd">
<day>05</day>
<month>01</month>
<year>2026</year>
</date>
<date date-type="accepted">
<day>23</day>
<month>01</month>
<year>2026</year>
</date>
</history>
<permissions>
<copyright-statement>Copyright &#x000A9; 2026 Nihngwo, Awazi and Kimengsi.</copyright-statement>
<copyright-year>2026</copyright-year>
<copyright-holder>Nihngwo, Awazi and Kimengsi</copyright-holder>
<license>
<ali:license_ref start_date="2026-02-19">https://creativecommons.org/licenses/by/4.0/</ali:license_ref>
<license-p>This is an open-access article distributed under the terms of the <ext-link ext-link-type="uri" xlink:href="https://creativecommons.org/licenses/by/4.0/">Creative Commons Attribution License (CC BY)</ext-link>. The use, distribution or reproduction in other forums is permitted, provided the original author(s) and the copyright owner(s) are credited and that the original publication in this journal is cited, in accordance with accepted academic practice. No use, distribution or reproduction is permitted which does not comply with these terms.</license-p>
</license>
</permissions>
<abstract>
<p>This study investigates how formal and informal institutions shape agroforestry processes and outcomes in the Western Highlands region of Cameroon. Using a mixed qualitative approach involving expert interviews, key informant interviews, and focus group discussions conducted in the Fokoue and Penka-Michel communities, the paper analyses the structure, processes, and effectiveness of formal (state-driven) and informal (community-based) institutions. Data were analyzed through descriptive statistics, chi-square tests, and thematic analysis. Results revealed a dualistic institutional system where formal institutions [comprising state agencies such as Ministry of Forestry and Wildlife (MINFOF), Ministry of Agriculture and Rural Development (MINADER), and Ministry of Livestock, Fisheries and Animal Husbandry (MINEPIA)] operate through codified laws, while informal institutions function through traditional councils, norms, and cultural taboos. Informal institutions demonstrated higher compliance and visibility within communities, largely due to social pressure, mutual support, and reverence for tradition. Conversely, formal institutions were perceived as ineffective because of weak policy frameworks and limited community engagement. Institutional effectiveness produced mixed outcomes: positive ecological effects (biodiversity conservation, soil improvement) and sociocultural cohesion through community networks, but also negative outcomes such as conflicts and weak coordination. The study concludes that agroforestry success is constrained by fragmented institutional collaboration and recommends strengthening policy coherence, integrating traditional governance systems into state frameworks, and promoting inclusive stakeholder partnerships to enhance sustainable agroforestry adoption and outcomes.</p></abstract>
<kwd-group>
<kwd>agroforestry institutions</kwd>
<kwd>Cameroon</kwd>
<kwd>community participation</kwd>
<kwd>governance</kwd>
<kwd>socio-ecological outcomes</kwd>
</kwd-group>
<funding-group>
  <funding-statement>The author(s) declared that financial support was not received for this work and/or its publication.</funding-statement>
</funding-group>
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<custom-meta-group>
<custom-meta>
<meta-name>section-at-acceptance</meta-name>
<meta-value>Natural Resources</meta-value>
</custom-meta>
</custom-meta-group>
</article-meta>
</front>
<body>
<sec sec-type="intro" id="s1">
<label>1</label>
<title>Introduction</title>
<p>Forest management institutions continue to attract global attention in the face of the rapid transformation of socioecological systems such as agroforestry. Agroforestry offers enhanced biodiversity conservation, higher overall productivity, provision of diverse ecosystem services, including water regulation and air quality improvement, and contributes significantly to sustainable development goals (<xref ref-type="bibr" rid="B42">Tebkew et al., 2024</xref>; <xref ref-type="bibr" rid="B4">Awazi et al., 2024a</xref>). Institutions are constructed as regulatory structures that provide meaning to social behavior, shaping human intervention (<xref ref-type="bibr" rid="B37">Scott, 1995</xref>). They manifest through cultures and structures in different contexts. To <xref ref-type="bibr" rid="B29">Ostrom (1990)</xref>, institutions are the set of rules in use by actors to organize activities that produce outcomes. The formality dichotomy presents institutions as formal (e.g., written and codified laws) and informal (e.g., rules that are unwritten or uncodified and are communicated through customs and traditions over generations) (<xref ref-type="bibr" rid="B46">Yeboah-Assiamah et al., 2017</xref>; <xref ref-type="bibr" rid="B15">Kimengsi et al., 2022</xref>).</p>
<p>Institutions are the outcome of choices determined by either exogenous or endogenous entities. Endogenous institutions are those which evolved with societies and are designed by the people that are likely to be affected by the designed structures or processes (<xref ref-type="bibr" rid="B38">Shvetsova, 2003</xref>; <xref ref-type="bibr" rid="B31">Owusu et al., 2023</xref>), while exogenous institutions are externally induced. For example, state rules introduced in communities assume exogenous, when compared to endogenous &#x02013; culturally embedded and largely informed rules. In this paper, institutions are defined using the structures-process approach (<xref ref-type="bibr" rid="B9">Fleetwood, 2008</xref>). Agroforestry-linked institutions consist of both informal village-based structures and formal or state organizations that enforce the rules and sanctions related to the access, use, and management of the forest. Additionally, they contain both informal (e.g., norms, values, taboos) and formal (laws) processes, which are provisioned to regulate the use of forest resources equally around protected areas. Both structures and processes are inseparable, as customs and beliefs evolve in the communities as agreed by traditional council. Furthermore, sanctions and rules determine the membership of such traditional institutions, thus making it difficult to separate structure from process (<xref ref-type="bibr" rid="B15">Kimengsi et al., 2022</xref>).</p>
<p>In Africa, agroforestry has taken center stage in scientific and policy discourses. From a systems perspective (<xref ref-type="bibr" rid="B22">Nair et al., 2021</xref>), agroforestry is defined as the deliberate integration of woody perennials (trees and shrubs) with crops and/or livestock on the same land-management unit, either spatially or temporally, where ecological and economic interactions occur among components. This definition emphasizes intentionality, interaction, and land-use integration. From a dynamic natural resource management perspective (<xref ref-type="bibr" rid="B19">Leakey, 1996</xref>), agroforestry is framed as a set of land-use systems and practices in which trees are deliberately managed alongside crops and/or animals to enhance productivity, sustainability, and livelihoods. This perspective emphasizes adaptability, multifunctionality, and socioecological processes. Although agroforestry practices and the farming system approach were introduced in the western region of Cameroon before the 1990s (<xref ref-type="bibr" rid="B26">Noubissie, 2020</xref>), the subsector has not been well-developed due to weak institutional support and poor incentive mechanisms. Today, Cameroon faces serious and linked problems: environmental degradation, population growth, and resource depletion. Furthermore, events in the colonial era resulted in land being unavailable to farmers, and thus people consider every inch of land to be a precious resource to be diligently safeguarded and meticulously managed. Additionally, despite the recognized benefits of agroforestry, its uptake has not been widespread due to challenges such as lack of support for agroforestry systems through public policies (<xref ref-type="bibr" rid="B5">Bishaw et al., 2013</xref>) and limited investment in the sector compared to intensified farming systems.</p>
<p>The scientific literature generally highlights the importance of collaboration between international and local actors to achieve the intended effects of agroforestry (<xref ref-type="bibr" rid="B2">Awazi, 2024</xref>; <xref ref-type="bibr" rid="B3">Awazi et al., 2024b</xref>). Several agroforestry interventions over the years have adopted a multifaceted approach by rolling out interventions in degraded and deforested landscapes with the involvement of diverse actors with various roles and interests. The literature also highlights the importance of institutions and the dynamics around institutions in different areas of forest management (in this case agroforestry) (<xref ref-type="bibr" rid="B15">Kimengsi et al., 2022</xref>; <xref ref-type="bibr" rid="B46">Yeboah-Assiamah et al., 2017</xref>). Currently, there are limited readily available data that provide the necessary information on these dimensions of agroforestry institutions, and an understanding of the role played by such types of formal or informal institutions in shaping agroforestry and the outcomes they produce is still missing. Hence this paper addresses the lacunae through three objectives. First, we aim to characterize the type of institutions in agroforestry. Second, we determine the effectiveness of the type (formal/informal) of institutions shaping agroforestry practices. Finally, we aim to evaluate the outcomes linked to the effectiveness of formal and informal institutions in agroforestry.</p></sec>
<sec id="s2">
<label>2</label>
<title>Analytical framework</title>
<p>The analytical framework of this paper centers on the formal and informal institution using an analytical lens (<xref ref-type="bibr" rid="B11">Ingram et al., 2015</xref>; <xref ref-type="bibr" rid="B15">Kimengsi et al., 2022</xref>). These institutions are typified as being either externally enforced (exogenous institutions) or internally enforced (self-enforcing, endogenous institutions) (<xref ref-type="bibr" rid="B10">Greif, 2006</xref>). In this regard, the elements of formality and informality could be identified in both exogenous and endogenous institutions (<xref ref-type="bibr" rid="B31">Owusu et al., 2023</xref>; <xref ref-type="bibr" rid="B28">Osei-Tutu et al., 2015</xref>; <xref ref-type="bibr" rid="B46">Yeboah-Assiamah et al., 2017</xref>).</p>
<p>Greater emphasis will be on their formality, that is, formal institutions&#x02014;documented and standardized structures and processes governed by the state (national authorities)&#x02014;and/or informal institutions&#x02014;uncodified practices with no government recognition which sometimes rely on unofficial arguments (<xref ref-type="bibr" rid="B46">Yeboah-Assiamah et al., 2017</xref>; <xref ref-type="bibr" rid="B31">Owusu et al., 2023</xref>). Both forms of institution (formal and informal) constantly interact to moderate the intentions and actions of resource users and actors, and hence shape the outcomes of agroforestry (<xref ref-type="fig" rid="F1">Figure 1</xref>). However, these institutions date as far back as the precolonial era and are rooted in customs and traditions in the communities (<xref ref-type="bibr" rid="B14">Kimengsi et al., 2023</xref>). Beliefs are linked to sacred forests and the practices of elders and traditional rulers in the communities.</p>
<fig position="float" id="F1">
<label>Figure 1</label>
<caption><p>Analytical framework of the study (adapted from <xref ref-type="bibr" rid="B15">Kimengsi et al., 2022</xref> and <xref ref-type="bibr" rid="B46">Yeboah-Assiamah et al., 2017</xref>).</p></caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fsrma-05-1732587-g0001.tif">
</graphic>
</fig>
<p>The outcomes of agroforestry interventions could be ecological-positive (e.g., tree planting, soil health improvements, biodiversity conservation, water regulations), ecological-negative (e.g., resistance to reforestation), economic-positive (e.g., increased income, improvement in local people&#x00027;s livelihood, diversification of income sources), economic-negative (e.g., market constraints, limited access to resources), social-positive (e.g., equity promotion, building of community resilience, and connection to the land), or social-negative (e.g., conflicts, frustrations of rural poor people) (see <xref ref-type="fig" rid="F1">Figure 1</xref>). These various outcomes are shaped by the different agroforestry systems. Some studies have explained that several interlinked outcomes (ecological, socioeconomic) shaped natural resource institutions (<xref ref-type="bibr" rid="B25">North, 1991</xref>). While formal institutions interact with informal ones, several interlinked eco-socio-economic interventions interact to shape institutional structures and processes. These agroforestry systems are reflected through the use of different practices such as home gardens, multipurpose trees, and shelterbelts (<xref ref-type="bibr" rid="B1">Awazi, 2022</xref>). This reflects the typology of institutions (<xref ref-type="fig" rid="F1">Figure 1</xref>) linked to the economic, ecological, and sociocultural benefits of agroforestry, which is likely to affect the overall outcomes of agroforestry.</p></sec>
<sec sec-type="materials and methods" id="s3">
<label>3</label>
<title>Materials and methods</title>
<sec>
<label>3.1</label>
<title>Study area</title>
<p>The Western Highlands, one of the five distinct agroecological zones in Cameroon and characterized by varying biophysical and climatic conditions, covers an area of approximately 3,807,280 hectares (38,072.8 square kilometers) (<xref ref-type="bibr" rid="B12">Jiotsa et al., 2015</xref>; <xref ref-type="bibr" rid="B45">WWF, 2022</xref>; <xref ref-type="bibr" rid="B20">MINFOF - MINEPDED, 2020</xref>). These highlands have a wavy topography characterized by savannah vegetation, stepped plateaus, low basins, hills, and plains intersected by gallery forests (<xref ref-type="bibr" rid="B20">MINFOF - MINEPDED, 2020</xref>). They contain one of the largest remaining patches of Afromontane forest in West Africa. The hydrographical network is dominated by streams and rivers. The climate is of tropical highland type characterized by two main seasons, rainy and dry. The average temperature is 25 &#x000B0;C, which is highly moderated by relief, which explains the usually cool nature of the entire region. The area has an altitude ranging between 1,000 and 2,500 meters above sea level (<xref ref-type="bibr" rid="B12">Jiotsa et al., 2015</xref>). The soils are ferralitic with high moisture retention (<xref ref-type="bibr" rid="B41">Tankou et al., 2017</xref>). Furthermore, it is one of the most densely populatedin Cameroon, with a population density of around 128.5 inhabitants per square kilometer (<xref ref-type="bibr" rid="B20">MINFOF - MINEPDED, 2020</xref>). This highland area comprises eight divisions (Menoua, Noun, Nde, Haut-Nkam, Mefi, Bamboutos, Haut-plateau, and Koung-khi).</p>
<p>The study area is situated in the Menoua division and has six subdivisions: Dschang, Santchou, Fongo-Tongo, Fokoue, Penka-Michel, and Nkong-Ni. Two of these subdivisions (Fokoue and Penka-Michel) are locations for the case studies in this article (<xref ref-type="fig" rid="F2">Figure 2</xref>). They have similar ecological, social, and economic characteristics (<xref ref-type="bibr" rid="B12">Jiotsa et al., 2015</xref>). Penka-Michel subdivision is geographically delimited to the north by Bamougoum, to the south by Bamendjou, and to the west by Kong-Nzem, and it has a surface area of 270 km<sup>2</sup> and a population of 75,182. It comprises of four groups: Baloum, Balessing, Bansoa, and Bamendou. Fokoue, in contrast, is made up of five groups: Bamegwou, Fotomenaq, Fotsa Touala, Fokoue, and Fomopea. It is delimited to the east by Batcham (Bamboutos) subdivisions, to the north by Foumbot, and to the west by Fongo-Tongo, and has a population of 11,053 (<xref ref-type="bibr" rid="B6">BUCREP, 2017</xref>). In this region, communities typically adhere to a traditional setup characterized by centralized structures and well-defined hierarchical institutions (<xref ref-type="bibr" rid="B12">Jiotsa et al., 2015</xref>; <xref ref-type="bibr" rid="B16">Kimengsi and Silberberger, 2023</xref>). The institutional setting of the communities in the region symbolizes a traditional African society where there is reverence for local customs and traditions.</p>
<fig position="float" id="F2">
<label>Figure 2</label>
<caption><p>Location of study communities in the West region of Cameroon.</p></caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fsrma-05-1732587-g0002.tif">
</graphic>
</fig>
<p>We therefore selected the Western Highlands of Cameroon for our study area because of (i) its ongoing land-use activities such as tree planting and agroforestry practices, (ii) the area&#x00027;s great need for agroforestry policies, and (iii) the well-structured institutional setup. Two rural communities (Fokoue and Penka-Michel) were selected from the area due to their similar socioeconomic activities and proximity to agroforestry.</p>
</sec>
<sec>
<label>3.2</label>
<title>Data collection</title>
<p>This paper forms part of a study on the institutional analysis of agroforestry processes in the Western Highlands region of Cameroon. Using thematic/narrative analysis, expert interviews, key informant interviews (KIIs), and focus group discussions (FGDs), we qualitatively explored the issues linked to the awareness of state provisions, policies, customs, and beliefs that influence agroforestry and the effectiveness of these institutions. The design of the research instruments (questionnaire and interview guides) and the data collection occurred in two communities of Cameroon&#x00027;s Western Highlands between June and July 2024. Research instruments (KII guide with eight questions, FGD guide capturing ten questions, and expert interviews guide comprising eight questions) all aided in data collection. Prior to the start of data collection, the research instruments were translated into French. We had English and French versions, which were reviewed by peers. Subsequently, the collection of information commenced. Seven KIIs were conducted in the study area, three in Fokoue communities and four in Penka-Michel communities, capturing information from traditional rulers of the communities, family heads, and forest users (<xref ref-type="fig" rid="F3">Figure 3</xref>). This was done to capture the perspective of almost all categories of people in the communities regarding the institutions that shape agroforestry practices, their effectiveness, and their outcomes. In addition to the eight questions that guided the KIIs, follow-up questions were asked to clarify the information provided by the respondents based on the responses received. On average, KIIs lasted 50 min.</p>
<fig position="float" id="F3">
<label>Figure 3</label>
<caption><p>Data collection framework.</p></caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fsrma-05-1732587-g0003.tif">
</graphic>
</fig>
<p>Additionally, using the FGD guide, five FGDs were conducted during the study. In each of the two study communities, FGDs took place with a mixed group (men, women, and youth), two groups of men, two of women, and one of youth. FGDs were done to obtain the perspective of all groups in the communities about the institutions that shape agroforestry practices and effectiveness. On average, FGDs comprised about six individuals and lasted 50 min. Finally, expert interviews were conducted to obtain experts&#x00027; perspectives on the formal and informal institutions that regulate rural Cameroon agroforestry: one senior divisional officer, one agriculture official, two forestry officials, one delegate of fisheries and animal husbandry, and one council official (mayor). The expert interview guide steered these interactions. As for the KIIs and FGDs, follow-up questions were asked to get further and better information on the subject. On average, expert interviews lasted 45 min. During data collection, questions and responses were predominantly in French.</p>
</sec>
<sec>
<label>3.3</label>
<title>Data analysis methods</title>
<p>Descriptive statistics were used to characterize the attributes of agroforestry-linked institutions in the two communities in the Western Highlands of Cameroon studied: the level of institutional arrangements (formal and informal) and the perception of agroforestry outcomes across ecological, economic, and social dimensions, that is, changes in the behavior of forest users. The variables used in the analysis are listed in <xref ref-type="table" rid="T1">Table 1</xref>. The dependent variables are the outcomes of agroforestry practices, and the independent variables are the attributes of the institutions&#x02014;awareness of state regulations, provisions, traditional customs, and community-based organizations.</p>
<table-wrap position="float" id="T1">
<label>Table 1</label>
<caption><p>List of variables and indicators used for data analysis.</p></caption>
<table frame="hsides" rules="groups">
<thead>
<tr>
<th valign="top" align="left"><bold>Dependent variables (outcomes)</bold></th>
<th valign="top" align="left"><bold>Independent variables (actor&#x02013;institution interaction)</bold></th>
</tr>
</thead>
<tbody>
<tr>
<td valign="top" align="left">&#x02022;Perception of increase in biodiversity<break/> &#x02022;Improvement in soil health<break/> &#x02022;Enhance market access<break/> &#x02022;Increase in income generation<break/> &#x02022;Social cohesion</td>
<td valign="top" align="left">&#x02022;Duration of agroforestry practice<break/> &#x02022;Types of agroforestry systems<break/> &#x02022;Government support<break/> &#x02022;Awareness of state regulations<break/> &#x02022;Awareness of community-based organizations<break/> &#x02022;Awareness of traditional customs<break/> &#x02022;Awareness of informal institutions</td>
</tr></tbody>
</table>
</table-wrap>
<p>The audio recorded during data collection was transcribed manually. Direct content and narrative analysis were used to analyze the data. The contents of the transcribed data were carefully studied in order to extract key information in line with the specific objectives of the study. Qualitative analysis software was not employed to assist with the analysis because issues related to institutions are complex and sometimes not directly framed, and so using software to assist with analysis might result in missing out certain key data. Our analysis focused on the roles of formal and informal agroforestry institutions, the outcomes of agroforestry institutions, and an exploration of the compliance levels of formal and informal institutions in the Western Highlands region of Cameroon.</p>
<p>For data processing, two types of analysis were differentiated according to the available qualitative and quantitative data. For quantitative data, the questionnaires were systematically input into a Microsoft Excel spreadsheet and analyzed using R. The R software was used to perform all statistical analysis (in particular, the chi-squared test) as the sample data were greater than 30 (a total of 103 households surveyed were sampled). The chi-squared test is a test of the independence of two variables, and it allowed us to conclude whether the associations observed deviated from what would be anticipated by chance. The predetermined probability threshold of 0.05 was considered. <italic>P</italic>-values that fell below 0.05 provided substantial evidence for the rejection of the null hypothesis of independence among the variables. Conversely, <italic>P</italic>-values that were greater than or equal to 0.05 failed to yield sufficient evidence for the rejection of the null hypothesis of independence among the variables. To augment the findings derived from the chi-squared test, a Cramer&#x00027;s V statistic was subsequently calculated to quantify the strength of (in)dependence between the variables. In terms of Cramer&#x00027;s V, values approaching 1 indicate a robust relationship, while a value nearing 0 signifies a weaker relationship between the variables of interest. However, values equal to or exceeding 3 are indicative of a strong dependence among the variables:</p>
<disp-formula id="E1"><mml:math id="M1"><mml:mtable columnalign="left"><mml:mtr><mml:mtd><mml:mrow><mml:mi>C</mml:mi><mml:mi>h</mml:mi><mml:mi>i</mml:mi><mml:mo>&#x02212;</mml:mo><mml:mi>s</mml:mi><mml:mi>q</mml:mi><mml:mi>u</mml:mi><mml:mi>a</mml:mi><mml:mi>r</mml:mi><mml:mi>e</mml:mi><mml:mi>d</mml:mi><mml:mo stretchy='false'>(</mml:mo><mml:msup><mml:mi>&#x003C7;</mml:mi><mml:mi>2</mml:mi></mml:msup><mml:mo stretchy='false'>)</mml:mo><mml:mo>=</mml:mo><mml:mstyle displaystyle="true"><mml:mo>&#x02211;</mml:mo></mml:mstyle><mml:mo stretchy='false'>(</mml:mo><mml:msup><mml:mrow><mml:mo stretchy='false'>(</mml:mo><mml:msub><mml:mi>O</mml:mi><mml:mi>i</mml:mi></mml:msub><mml:mo>&#x02212;</mml:mo><mml:msub><mml:mi>E</mml:mi><mml:mi>i</mml:mi></mml:msub><mml:mo stretchy='false'>)</mml:mo></mml:mrow><mml:mn>2</mml:mn></mml:msup><mml:mo>/</mml:mo><mml:msub><mml:mi>E</mml:mi><mml:mi>i</mml:mi></mml:msub><mml:mo stretchy='false'>)</mml:mo><mml:mo>,</mml:mo></mml:mrow></mml:mtd></mml:mtr></mml:mtable></mml:math></disp-formula>
<p>where <italic>O</italic><sub><italic>i</italic></sub> is the observed frequency of the variables, <italic>E</italic><sub><italic>i</italic></sub> is the expected frequency of the variables, &#x003C7;<sup>2</sup> is the chi-squared statistic, and n is the number of observations.</p>
<p>For qualitative data, our analysis was conducted through narratives and thematic analyses relating to the knowledge and practice of agroforestry, related customs and traditions, and its effectiveness and outcomes. These narratives provided a rich insight into the lived experience of institutions in agroforestry.</p></sec>
</sec>
<sec sec-type="results" id="s4">
<label>4</label>
<title>Results</title>
<sec>
<label>4.1</label>
<title>Characterization of formal and informal agroforestry institutions</title>
<p>Our research revealed a dualistic institutional framework. These institutions are linked to formal (exogenous) and informal (endogenous) structures and processes (<xref ref-type="fig" rid="F4">Figure 4</xref>). The formal institutions are state-driven entities, operating through codified policies (e.g., the 1994 Forestry Law recently updated to the 2024 Forestry and Wildlife Law). These include the government ministries&#x02014;MINADER, MINFOF, and MINEPIA&#x02014;alongside local government authorities (senior divisional officer, mayor). These bodies are mandated with the top-down implementation of national policies, provision of technical support, and enforcement of land-use regulations. However, the study revealed an absence of a de jure policy (no specific formal laws) tailored to agroforestry, resulting in the <italic>ad hoc</italic> application of general agricultural and forestry regulations.</p>
<fig position="float" id="F4">
<label>Figure 4</label>
<caption><p>Typologies of institutions that shape agroforestry in the WHC.</p></caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fsrma-05-1732587-g0004.tif">
</graphic>
</fig>
<p>Conversely, informal institutions are endogenous community-based systems of governance rooted in local customs and traditional ecological knowledge. These institutions&#x02014;traditional councils&#x02014;are led by chiefs and elders who act as custodians of cultural heritage and arbiters of land-related disputes. Key structures facilitating collective action are njangi groups (traditional rotating savings and credit associations), which promote resource pooling, labor sharing, and knowledge exchange. However, the processes governing these institutions are unwritten but socially enforced; for example, sacred days &#x0201C;njang, mete&#x0201D; prohibit farming or tree cutting, certain trees (e.g., the baobab) are protected for cultural reasons, and trees are planted to mark significant life events (e.g., childbirth). Of these institutions, the most dominant are informal community-based organizations, particularly the njangi (loan and thrift) groups, which have more visible influence and participation (50%) compared to formal institutions, where role is seen as largely ineffective with low awareness among farmers (43%) but exhibit fragmentation and limit collaboration at local level. Additionally, although formal institutions provide national policies and rules, the actual practices of agroforestry are largely rooted in traditions and community mutual support groups.</p>
</sec>
<sec>
<label>4.2</label>
<title>Effectiveness/compliance levels of formal and informal agroforestry institutions</title>
<p>From the evidence of our study, a significant difference was observed between the perceived effectiveness of formal and informal institutions. Formal state-led institutions were largely considered ineffective by local communities. The research revealed that a substantial proportion of respondents (43.7%) were unaware of government policies or support pertaining to agroforestry pertinent to agroforestry, and a majority (59.2%) indicated no direct engagement with these institutions. A small proportion of respondents rated them as &#x0201C;somewhat effective&#x0201D; (6%) or &#x0201C;very effective&#x0201D; (4%), while 15% deemed them not effective at all. This perceived inefficacy is linked to inadequate communication, inconsistent implementation of state-led programmes, and enforcement mechanisms perceived as punitive, such as fines for tree cutting, which act as a discouragement to farmers&#x00027; participation. A female FGD participant recounted: &#x0201C;We currently receive no support from the government, leaving each individual to explore on their own without formal guidelines to follow. We rely primarily on our existing knowledge and resources, and our methods of practice here remain largely traditional.&#x0201D;</p>
<p>This supports the statement from the agriculture delegate:</p>
<disp-quote><p>&#x0201C;There are no formal laws or regulations impacting or influencing agroforestry. While farming is widely practiced, farmers often grow fruit trees in their farms and dwellings alongside crops. There are no specific regulations that guide agroforestry in the community. However, there is a traditional guideline requiring everyone to plant at least one avocado tree on their land.&#x0201D;</p></disp-quote>
<p>However, the informal institutions demonstrated greater salience and influence within the community. Just over half of the respondents (50.5%) actively participated in community-based organizations. Compliance with informal rules is high and is enforced through mechanisms of social pressure, mutual reciprocity, and adherence to cultural traditions. While these endogenous institutions are effective in fostering local collaboration and upholding traditional conservation practices, their capacity to scale agroforestry initiatives up is constrained by a lack of formal recognition and access to external resources.</p>
<p>With respect to the effectiveness/compliance level of the informal institutions, our study revealed a high degree of institutional visibility and active community participation as primary drivers of their effectiveness. For instance, 50% were actively involved in community groups such as agricultural cooperatives and njangi associations. This level of engagement facilitates robust networks for knowledge dissemination, resource sharing, and collective action, establishing these institutions as a vehicle for local agroforestry. Additionally, it was reported that many people in the communities have reverence for traditional customary practices related to the environment&#x02014;such as maintaining trees for cultural rituals and cultivating fruit trees inherited from their ancestors&#x02014;and, by extension, agroforestry is related to the community&#x00027;s belief system relating to the protection of sacred trees/forests; this is in preference to punitive, top-down enforcement, which is often perceived as coercive and consequently is less effective. A male FGD participant recounted:</p>
<disp-quote><p>&#x0201C;In our community, there are no specific rules or customs guiding agroforestry practices. While agroforestry is practiced, it often occurs without deep understanding and is done traditionally However, we observe certain days, like &#x0201C;Bowang,&#x0201D; &#x0201C;njang,&#x0201D; &#x0201C;ngang,&#x0201D; when no work is allowed. We are aware of these days and respect them. Additionally, there are sacred places surrounded by small forests. These areas are considered untouchable, and no one is allowed to cut trees or disturb them. These sacred forests are located near the chiefdom, and everyone in the village, even children, understands and respects these boundaries.&#x0201D;</p></disp-quote>
<p>This supports the information from a notable of the Bansoa community:</p>
<disp-quote><p>&#x0201C;In the tradition, certain trees like the baobab (maboku) are considered sacred and are never cut down. These trees are seen as protectors of the chiefdom. These trees grow naturally without being planted. Other plants, such as lianas, are also preserved within chiefdoms. The presence of a baobab tree often signifies the location of a chiefdom.&#x0201D;</p></disp-quote>
</sec>
<sec>
<label>4.3</label>
<title>Outcomes of agroforestry institutional effectiveness/compliance</title>
<p>Agroforestry institutions in the Western Highlands of Cameroon produce positive and negative ecological, economic, and sociocultural outcomes (<xref ref-type="fig" rid="F5">Figure 5</xref>). While formal institutions produce ecological, economic, and sociocultural outcomes, informal institutions produce positive ecological outcomes and positive and negative economic and sociocultural outcomes.</p>
<fig position="float" id="F5">
<label>Figure 5</label>
<caption><p>Socioeconomic benefits.</p></caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fsrma-05-1732587-g0005.tif">
</graphic>
</fig>
<p>For the informal institutions, the research revealed a positive ecological link to informal practices such as the integration of fruit trees, for example, avocado (common in the Bamileke tradition), that support pollinators, which are statistically proven to be associated with perceived increased in biodiversity (<italic>p</italic> &#x0003C; 0.001; Cramer&#x00027;s V = 0.419). This is achieved through the preservation and protection of sacred forests, which act as de facto nature reserves. These advantages are mitigated by unsustainable practices including the destruction of raffia palms for market gardening and the excess use of chemical fertilizers. While negative ecological outcomes were observed in formal institutions, they fail to provide a coherent agroforestry policy. The absence of regulations and oversight contributes to persistent deforestation and the widespread cultivation of eucalyptus monocultures, which are associated with soil nutrient depletion. The actions of some formal structures, such as the municipal council authorizing tree felling, cause biodiversity loss and habitat destruction.</p>
<p>The socioeconomic benefits perceived by farmers (<xref ref-type="fig" rid="F5">Figure 5</xref>) were livelihood improvement (50%) through diversified products for subsistence and local sale. Through structures such as njangi groups, farmers could pool resources and labor. However, the negative economic outcome is the failure to scale. While effective at the local level, the informal institutions lack the formal recognition and capacity to develop value chains or connect farmers to bigger markets. Thus, they remain confined to subsistence and small-scale enterprises. Additionally, the negative economic outcome associated with formal institutions is the shifting of potential. Failure to provide support, especially in developing market access, was identified as a blockade to the expansion of economic benefits. The historical abandonment of coffee farming, a key economic activity cash crop, has contributed to youth out-migration (rural exodus):</p>
<disp-quote><p>&#x0201C;We don&#x00027;t know what the state&#x00027;s provisions are in terms of agroforestry. In terms of agricultural cultivation, we heard of fertilisers that were subsidised over the years, but we the Bamendou growers haven&#x00027;t benefited, we&#x00027;ve been abandoned. As for rules and policies in agroforestry, we are free to plant as we like, but the major problem is that there&#x00027;s no follow-up.&#x0201D;</p></disp-quote>
<p>Also, as recounted by a forest technician and head of an NGO (SAFEWORK),</p>
<disp-quote><p>&#x0201C;Currently, targeted subsidies for agroforestry are not available. Even coffee farming, once a highly developed agroforestry practice, has largely disappeared due to economic challenges. The global market price of coffee has sharply declined, and the government has not intervened to stabilise or support local prices.&#x0201D;</p></disp-quote>
<p>Regarding the social outcome of agroforestry for formal institutions, the study revealed a significant source of social conflict and tension. Their top-down, punitive enforcement rules, such as fines on farmers for cutting trees without permits, is perceived as repressive and weakens trust between the state and the communities. This approach creates resentment and discourages farmer participation. Also, the conflicts between the conservation goals of MINFOF and MINADER and the revenue generation priorities of the local council, creates confusion and aggravates tensions, weakening the overall governance fabrics. Furthermore, the analysis shows the association between community-based organizations and formal institutions to be weak (&#x003C7;<sup>2</sup>, <italic>p</italic> = 0.685, Cramer&#x00027;s V = 0.093). This could be because engaging in a community organization does not necessarily correspond with increased knowledge of formal institutions, and thus collaboration can be fragmented. While the negative social outcomes were more dominant in formal institutions, the key positive social outcomes associated with informal institutions include community wellbeing and social cohesion. This is through mechanisms of mutual support, knowledge exchange, and collective action. By upholding traditions and resolving disputes internally, they reinforce cultural identity and strengthen community bonds. However, the social resilience they build is vulnerable. The study reported farmers&#x00027; intent to establish agroforestry groups, but a decline of more formal farmer groups (CIGs) due to mistrust and mismanagement shows that even community-based structures can be fragile. Additionally, the social harmony they foster is consistently threatened by conflicts arising from the actions of formal institutions, demonstrating their limited power to shield the community from external governance failures.</p></sec>
</sec>
<sec sec-type="discussion" id="s5">
<label>5</label>
<title>Discussion</title>
<sec>
<label>5.1</label>
<title>Characterization of formal and informal agroforestry institutions</title>
<p>Our results revealed that a diverse range of formal and informal institutions shape agroforestry in the Western Highlands of Cameroon. The key formal structure includes government ministries (MINFOF, MINADER, MINEPIA), which operate through codified legal laws such as the 1994 Forestry Law (updated to the 2024 Forestry and Wildlife Law), and the informal institutions are rooted in local customs, norms, and traditional knowledge, embodied by structures including traditional councils, youth groups, and njangi (loan and thrift) groups. However, the informal structures do not denote an entire setting as some traditional councils are recognized by administrative authorities, and they are allowed to perform during festivals. This characterization aligns with established theories of polycentric governance, where multiple, overlapping centers of authority govern resource use (<xref ref-type="bibr" rid="B30">Ostrom, 2010</xref>). However, our findings indicated that these polycentric systems operate separately and are not well-connected. There is a significant gap between the formal institutions and the actual day-to-day rules that the community follow (informal). Our findings align with recent research in other sub-Saharan African contexts, which highlights that state-led, top-down structures often fail to connect with local realities, leaving informal institutions to fill the void (<xref ref-type="bibr" rid="B21">Mukong et al., 2025</xref>; <xref ref-type="bibr" rid="B15">Kimengsi et al., 2022</xref>). Additionally, a key finding is the absence of specific, clear policies for agroforestry in rural Cameroon; existing farming and forestry laws are used instead but are often fragmented and do not operate well. However, our results differ from the findings in parts of Southeast Asia that reported clear agroforestry policies implemented to improve coordination and effectiveness (<xref ref-type="bibr" rid="B24">Ndlovu, 2025</xref>; <xref ref-type="bibr" rid="B43">Venn et al., 2024</xref>). In addition, the result is inconsistent with the polycentric governance model (<xref ref-type="bibr" rid="B13">Katic, 2021</xref>), where multiple stakeholders operate across scales. The dominance of informal institutions in Cameroon&#x00027;s agroforestry plays a significant force in managing agroforestry in the region. Hence, effective collaboration between the formal and informal institutions, with strong partnership and alignment goals and strategies, is crucial to enhance effective agroforestry interventions.</p>
</sec>
<sec>
<label>5.2</label>
<title>Effectiveness/compliance levels of formal and informal agroforestry institutions</title>
<p>Based on our study, we found that the effectiveness of institutional typologies in agroforestry is different. The results shed light on the dynamics of effectiveness with institutions within the context of agroforestry in the Western Highlands of Cameroon. The observed disparities in effectiveness bring a nuanced understanding of how local communities and formal entities engage with and respond to different institutions in the context of agroforestry. In addition to formal institutions, the absence of targeted agroforestry policies contributed to a low awareness of government initiatives by communities and limited participation with little direct engagement. This is often rooted in the inadequate commitment and self-interest of officials responsible for enforcing these institutions. Regulations are often perceived as harsh or punitive through fines for unauthorized tree cutting or livestock issues, which discourages farmer participation. There is, therefore, a need to ameliorate the enforcement mechanism of the formal agroforestry-related institutions and their integration with informal systems. This result is similar to that of other studies with non-compliance with formal institutions (<xref ref-type="bibr" rid="B48">Zouari and Hachicha, 2024</xref>; <xref ref-type="bibr" rid="B32">Owusu et al., 2024</xref>).</p>
<p>In contrast, the effectiveness associated with informal institutions could be due to high visibility and acceptance within communities. Their rules are upheld via social pressures, reciprocity, and respect for traditions. Our results can be attributed to rural communities&#x00027; absolute reverence for their culture, which is closely linked to norms and traditions in honoring sacred days and protecting culturally valued trees. These mechanisms provide effective, non-coercive governance on environmental management (<xref ref-type="bibr" rid="B21">Mukong et al., 2025</xref>). However, informal institutions lack formal recognition and access to external resources, limiting their ability to expand agroforestry beyond local or subsistence levels. This gap highlights the need to better integrate informal institutions into the formal governance framework to enable their broader contribution (<xref ref-type="bibr" rid="B47">Zinngrebe et al., 2020</xref>). This result corroborates the findings of <xref ref-type="bibr" rid="B16">Kimengsi and Silberberger (2023)</xref>, who held that the actors driving exogenous institutions in the context of climate adaptation in rural Cameroon should accommodate endogenous cultural institutional arrangements and vice versa. Furthermore, there was a pronounced adherence of formal and informal institutions (the planting of avocado) that exhibits a distinct characteristic of effectiveness that cannot be underestimated. While this result holds for the Western Highlands, it cannot be extrapolated for the different agroecological zones of Cameroon, where variations in the effectiveness of informal institutions are expected. This is linked to the ethnic diversity of Cameroon, which contains more than 250 ethnic groups (<xref ref-type="bibr" rid="B8">Fearon, 2003</xref>), implying significant variations in the informal cultural institutions that either enhance or constrain compliance (<xref ref-type="bibr" rid="B27">Nuesiri, 2012</xref>). For instance, in the Greater South region, significant variations in compliance with endogenous cultural institutions are observed (<xref ref-type="bibr" rid="B14">Kimengsi et al., 2023</xref>). Therefore, the findings from this landscape (the Western Highlands) cannot be used to represent the whole of Cameroon or sub-Saharan Africa. It is imperative that additional studies be conducted in other parts of the region to assist in providing a holistic picture of the compliance level of agroforestry-related institutions.</p>
</sec>
<sec>
<label>5.3</label>
<title>Outcomes of agroforestry linked to institutional effectiveness</title>
<p>Our study revealed that the effectiveness of informal institutions is linked to positive ecological outcomes. This could be due to the local traditions that preserve sacred forest and fruit tree cultivation, benefiting biodiversity, especially pollinators. The analysis of the study revealed that there is an association between awareness of formal institutions and improvement in on-farm biodiversity and soil health, which is achieved through the maintenance of traditional ecological knowledge. This finding corroborates other studies on the role of communit- based institutions in enhancing biodiversity within agroforestry systems (<xref ref-type="bibr" rid="B17">Kuyah et al., 2022</xref>; <xref ref-type="bibr" rid="B42">Tebkew et al., 2024</xref>).</p>
<p>In contrast, the low compliance of formal institutions is directly linked to negative ecological outcomes. The failure to regulate land use is a result of fragmented and weak governance. This could be due to persistent deforestation and an increase in eucalyptus monocultures. Our result aligns with studies in tropical regions that reported monoculture deplete nutrients and reduce ecosystem services (<xref ref-type="bibr" rid="B7">Farley et al., 2005</xref>; <xref ref-type="bibr" rid="B44">Vigo et al., 2025</xref>). However, our results were inconsistent with studies from regions with strong state-led conservation programmes where formal enforcement has been shown to curb deforestation (<xref ref-type="bibr" rid="B36">Sarmiento et al., 2024</xref>).</p>
<p>Regarding informal institutions, key positive economic and social outcomes are rooted in improved livelihoods through knowledge sharing and participation. However, the negative outcome is their inability to scale agroforestry products due to limited capacity to access markets. With limited formal recognition and provisions, their economic impact remains localized leading to abandoned cash crops (e.g., coffee), causing rural youth to migrate for work. This study resonates with the research of <xref ref-type="bibr" rid="B18">Lawasi et al. (2024)</xref>, who reported that limited market access directly affected farmers&#x00027; ability to profit from sustainable farming practices, but diverges from that of <xref ref-type="bibr" rid="B33">Paudel et al. (2022)</xref> in Nepal, who reported a stronger market-driven adoption. However, another positive sociocultural outcome is the spiritual connection of the local people to their land and their ancestral spirits. For example, some studies reported that forest restoration and protection via sacred forests connects local people to their land and ancestors (<xref ref-type="bibr" rid="B40">Sullivan et al., 2024</xref>; <xref ref-type="bibr" rid="B39">Sinthumule, 2024</xref>). However, social stability faces threats from declining formal farmer groups due to mistrust and ineffective management, and from resentments caused by punitive actions by formal authorities. Such tensions undermine trust and weaken inclusive governance needed for sustainable agroforestry (<xref ref-type="bibr" rid="B23">Ndiovu et al., 2025</xref>). Our research is consistent with the findings of <xref ref-type="bibr" rid="B35">Reed et al. (2024)</xref>.</p>
<p>Regarding formal institutions, limited provisions and an unawareness of state provisions linked to agroforestry is rooted in the absence of support for market access and follow-up that prevents the scaling of agroforestry from a subsistence activity to a viable economic enterprise. Moreover, mistrust and conflicts discourage farmer participation in sustainable practices. Our research resonates with findings by <xref ref-type="bibr" rid="B34">Puspita et al. (2024)</xref>, who argue that government intervention, if too controlling, can erode or destroy community social capital, thereby weakening collective action in water resource management settings.</p></sec>
</sec>
<sec sec-type="conclusion" id="s6">
<label>6</label>
<title>Conclusion</title>
<p>Governance of agroforestry has become a subject of interest among scientist and policy makers. Our study used the formal and informal institutional analytical lens to characterize formal and informal institutions, their effectiveness/compliance levels, and the outcomes linked to the effectiveness of these institutions in agroforestry. We observed that both formal (state regulations, policies) and informal (customs, traditions) institutions are crucial in governing agroforestry. While formal institutions provide a structured framework, informal institutions deeply embedded in local culture often dictate the practical application and compliance within local communities. Awareness of and adherence to formal institutions were low compared to the high compliance observed with informal community-driven structures. This implies that cultural reverence for traditional norms significantly enhances the effectiveness of informal institutions in shaping agroforestry practices. Finally, positive outcomes, such as tree planting, that increase income, diversify livelihoods, and aid community resilience are more pronounced when agroforestry aligns with existing informal institutions. Conversely, a sole reliance on formal instructions without considering local customs could lead to suboptimal outcomes such as resistance to reforestation and social friction. This study therefore highlights the importance of integrating traditional knowledge and practices into future agroforestry policies to promote legal compliance and more sustainable and equitable outcomes in similar socioecological context.</p></sec>
</body>
<back>
<sec sec-type="data-availability" id="s7">
<title>Data availability statement</title>
<p>The raw data supporting the conclusions of this article will be made available by the authors, without undue reservation.</p>
</sec>
<sec sec-type="ethics-statement" id="s8">
<title>Ethics statement</title>
<p>The studies involving humans were approved by Faculty of Environmental Sciences, Technische Universitat Dresden, Tharandt, Germany. The studies were conducted in accordance with the local legislation and institutional requirements. The participants provided their written informed consent to participate in this study.</p>
</sec>
<sec sec-type="author-contributions" id="s9">
<title>Author contributions</title>
<p>AN: Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Project administration, Resources, Software, Validation, Visualization, Writing &#x02013; original draft, Writing &#x02013; review &#x00026; editing. NA: Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Project administration, Resources, Software, Supervision, Validation, Visualization, Writing &#x02013; original draft, Writing &#x02013; review &#x00026; editing. JK: Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Project administration, Resources, Software, Supervision, Validation, Visualization, Writing &#x02013; original draft, Writing &#x02013; review &#x00026; editing.</p>
</sec>
<sec sec-type="COI-statement" id="conf1">
<title>Conflict of interest</title>
<p>The author(s) declared that this work was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest.</p>
<p>The authors NA and JK declared that they were an editorial board member of Frontiers, at the time of submission. This had no impact on the peer review process and the final decision.</p>
</sec>
<sec sec-type="ai-statement" id="s11">
<title>Generative AI statement</title>
<p>The author(s) declared that generative AI was not used in the creation of this manuscript.</p>
<p>Any alternative text (alt text) provided alongside figures in this article has been generated by Frontiers with the support of artificial intelligence and reasonable efforts have been made to ensure accuracy, including review by the authors wherever possible. If you identify any issues, please contact us.</p></sec>
<sec sec-type="disclaimer" id="s12">
<title>Publisher&#x00027;s note</title>
<p>All claims expressed in this article are solely those of the authors and do not necessarily represent those of their affiliated organizations, or those of the publisher, the editors and the reviewers. Any product that may be evaluated in this article, or claim that may be made by its manufacturer, is not guaranteed or endorsed by the publisher.</p>
</sec>
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<fn fn-type="custom" custom-type="edited-by" id="fn0001">
<p>Edited by: <ext-link ext-link-type="uri" xlink:href="https://loop.frontiersin.org/people/935004/overview">Joseph George Ray</ext-link>, Mahatma Gandhi University, India</p>
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<fn fn-type="custom" custom-type="reviewed-by" id="fn0002">
<p>Reviewed by: <ext-link ext-link-type="uri" xlink:href="https://loop.frontiersin.org/people/572841/overview">Verina Jane Ingram</ext-link>, Wageningen University and Research, Netherlands</p>
<p><ext-link ext-link-type="uri" xlink:href="https://loop.frontiersin.org/people/2622806/overview">Vellayati Hajad</ext-link>, Universitas Teuku Umar, Indonesia</p>
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