<?xml version="1.0" encoding="UTF-8"?>
<!DOCTYPE article PUBLIC "-//NLM//DTD JATS (Z39.96) Journal Publishing DTD v1.3 20210610//EN" "JATS-journalpublishing1-3-mathml3.dtd">
<article xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:ali="http://www.niso.org/schemas/ali/1.0/" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance" article-type="research-article" dtd-version="1.3" xml:lang="EN">
<front>
<journal-meta>
<journal-id journal-id-type="publisher-id">Front. Mar. Sci.</journal-id>
<journal-title-group>
<journal-title>Frontiers in Marine Science</journal-title>
<abbrev-journal-title abbrev-type="pubmed">Front. Mar. Sci.</abbrev-journal-title>
</journal-title-group>
<issn pub-type="epub">2296-7745</issn>
<publisher>
<publisher-name>Frontiers Media S.A.</publisher-name>
</publisher>
</journal-meta>
<article-meta>
<article-id pub-id-type="doi">10.3389/fmars.2025.1737039</article-id>
<article-version article-version-type="Version of Record" vocab="NISO-RP-8-2008"/>
<article-categories>
<subj-group subj-group-type="heading">
<subject>Original Research</subject>
</subj-group>
</article-categories>
<title-group>
<article-title>Disentangling Eastern Pacific Warming: El Ni&#xf1;o 2023&#x2013;2024 <italic>vs</italic> seasonal Panam&#xe1; Bay influence</article-title>
</title-group>
<contrib-group>
<contrib contrib-type="author" corresp="yes">
<name><surname>Ormaza-Gonz&#xe1;lez</surname><given-names>Franklin Isaac</given-names></name>
<xref ref-type="aff" rid="aff1"><sup>1</sup></xref>
<xref ref-type="corresp" rid="c001"><sup>*</sup></xref>
<uri xlink:href="https://loop.frontiersin.org/people/963464/overview"/>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="conceptualization" vocab-term-identifier="https://credit.niso.org/contributor-roles/conceptualization/">Conceptualization</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="methodology" vocab-term-identifier="https://credit.niso.org/contributor-roles/methodology/">Methodology</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Formal analysis" vocab-term-identifier="https://credit.niso.org/contributor-roles/formal-analysis/">Formal analysis</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Writing &#x2013; review &amp; editing" vocab-term-identifier="https://credit.niso.org/contributor-roles/writing-review-editing/">Writing &#x2013; review &amp; editing</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="supervision" vocab-term-identifier="https://credit.niso.org/contributor-roles/supervision/">Supervision</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="validation" vocab-term-identifier="https://credit.niso.org/contributor-roles/validation/">Validation</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Writing &#x2013; original draft" vocab-term-identifier="https://credit.niso.org/contributor-roles/writing-original-draft/">Writing &#x2013; original draft</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="investigation" vocab-term-identifier="https://credit.niso.org/contributor-roles/investigation/">Investigation</role>
</contrib>
<contrib contrib-type="author">
<name><surname>Martillo-Bustamante</surname><given-names>Carlos</given-names></name>
<xref ref-type="aff" rid="aff1"><sup>1</sup></xref>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="visualization" vocab-term-identifier="https://credit.niso.org/contributor-roles/visualization/">Visualization</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Data curation" vocab-term-identifier="https://credit.niso.org/contributor-roles/data-curation/">Data curation</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="software" vocab-term-identifier="https://credit.niso.org/contributor-roles/software/">Software</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="resources" vocab-term-identifier="https://credit.niso.org/contributor-roles/resources/">Resources</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Writing &#x2013; review &amp; editing" vocab-term-identifier="https://credit.niso.org/contributor-roles/writing-review-editing/">Writing &#x2013; review &amp; editing</role>
</contrib>
<contrib contrib-type="author">
<name><surname>Espinoza</surname><given-names>Elvis</given-names></name>
<xref ref-type="aff" rid="aff1"><sup>1</sup></xref>
<uri xlink:href="https://loop.frontiersin.org/people/2895170/overview"/>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Writing &#x2013; review &amp; editing" vocab-term-identifier="https://credit.niso.org/contributor-roles/writing-review-editing/">Writing &#x2013; review &amp; editing</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="software" vocab-term-identifier="https://credit.niso.org/contributor-roles/software/">Software</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Data curation" vocab-term-identifier="https://credit.niso.org/contributor-roles/data-curation/">Data curation</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="visualization" vocab-term-identifier="https://credit.niso.org/contributor-roles/visualization/">Visualization</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Formal analysis" vocab-term-identifier="https://credit.niso.org/contributor-roles/formal-analysis/">Formal analysis</role>
</contrib>
<contrib contrib-type="author">
<name><surname>Proust</surname><given-names>Jean-Noel</given-names></name>
<xref ref-type="aff" rid="aff2"><sup>2</sup></xref>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Writing &#x2013; review &amp; editing" vocab-term-identifier="https://credit.niso.org/contributor-roles/writing-review-editing/">Writing &#x2013; review &amp; editing</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="resources" vocab-term-identifier="https://credit.niso.org/contributor-roles/resources/">Resources</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Funding acquisition" vocab-term-identifier="https://credit.niso.org/contributor-roles/funding-acquisition/">Funding acquisition</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Project-administration" vocab-term-identifier="https://credit.niso.org/contributor-roles/project-administration/">Project administration</role>
</contrib>
<contrib contrib-type="author">
<name><surname>Michaud</surname><given-names>Fran&#xe7;ois</given-names></name>
<xref ref-type="aff" rid="aff3"><sup>3</sup></xref>
<xref ref-type="aff" rid="aff4"><sup>4</sup></xref>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="supervision" vocab-term-identifier="https://credit.niso.org/contributor-roles/supervision/">Supervision</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Writing &#x2013; review &amp; editing" vocab-term-identifier="https://credit.niso.org/contributor-roles/writing-review-editing/">Writing &#x2013; review &amp; editing</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="resources" vocab-term-identifier="https://credit.niso.org/contributor-roles/resources/">Resources</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Funding acquisition" vocab-term-identifier="https://credit.niso.org/contributor-roles/funding-acquisition/">Funding acquisition</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Project-administration" vocab-term-identifier="https://credit.niso.org/contributor-roles/project-administration/">Project administration</role>
</contrib>
<contrib contrib-type="author">
<name><surname>Lebrun</surname><given-names>Jean-Frederic</given-names></name>
<xref ref-type="aff" rid="aff5"><sup>5</sup></xref>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Data curation" vocab-term-identifier="https://credit.niso.org/contributor-roles/data-curation/">Data curation</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Formal analysis" vocab-term-identifier="https://credit.niso.org/contributor-roles/formal-analysis/">Formal analysis</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="software" vocab-term-identifier="https://credit.niso.org/contributor-roles/software/">Software</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="resources" vocab-term-identifier="https://credit.niso.org/contributor-roles/resources/">Resources</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="investigation" vocab-term-identifier="https://credit.niso.org/contributor-roles/investigation/">Investigation</role>
<role vocab="credit" vocab-identifier="https://credit.niso.org/" vocab-term="Writing &#x2013; review &amp; editing" vocab-term-identifier="https://credit.niso.org/contributor-roles/writing-review-editing/">Writing &#x2013; review &amp; editing</role>
</contrib>
</contrib-group>
<aff id="aff1"><label>1</label><institution>Escuela Superior Polit&#xe9;cnica del Litoral (ESPOL), Facultad de Ingenier&#xed;a Mar&#xed;tima y Ciencias del Mar, Polytechnic University</institution>, <city>Guayaquil</city>,&#xa0;<country country="ec">Ecuador</country></aff>
<aff id="aff2"><label>2</label><institution>Univ Rennes, French National Centre for Scientific Research (CNRS), G&#xe9;osciences Rennes</institution>, <city>Rennes</city>,&#xa0;<country country="fr">France</country></aff>
<aff id="aff3"><label>3</label><institution>Facult&#xe9; des Sciences et Ing&#xe9;nierie, Sorbonne Universit&#xe9;</institution>, <city>Paris</city>,&#xa0;<country country="fr">France</country></aff>
<aff id="aff4"><label>4</label><institution>Universit&#xe9; C&#xf4;te d&#x2019;Azur, French National Research Institute for Sustainable Development (IRD), French National Centre for Scientific Research (CNRS), Observatoire de la C&#xf4;te d&#x2019;Azur, Laboratoire G&#xe9;oazur</institution>, <city>Valbonne</city>,&#xa0;<country country="fr">France</country></aff>
<aff id="aff5"><label>5</label><institution>Geosciences Montpellier, Universite des Antilles, French National Centre for Scientific Research (CNRS), Universite de Montpellier</institution>, <city>Pointe a Pitre</city>,&#xa0;<country country="gp">Guadeloupe</country></aff>
<author-notes>
<corresp id="c001"><label>*</label>Correspondence: Franklin Isaac Ormaza-Gonz&#xe1;lez, <email xlink:href="mailto:formaza@espol.edu.ec">formaza@espol.edu.ec</email></corresp>
</author-notes>
<pub-date publication-format="electronic" date-type="pub" iso-8601-date="2026-01-19">
<day>19</day>
<month>01</month>
<year>2026</year>
</pub-date>
<pub-date publication-format="electronic" date-type="collection">
<year>2025</year>
</pub-date>
<volume>12</volume>
<elocation-id>1737039</elocation-id>
<history>
<date date-type="received">
<day>31</day>
<month>10</month>
<year>2025</year>
</date>
<date date-type="accepted">
<day>09</day>
<month>12</month>
<year>2025</year>
</date>
<date date-type="rev-recd">
<day>08</day>
<month>12</month>
<year>2025</year>
</date>
</history>
<permissions>
<copyright-statement>Copyright &#xa9; 2026 Ormaza-Gonz&#xe1;lez, Martillo-Bustamante, Espinoza, Proust, Michaud and Lebrun.</copyright-statement>
<copyright-year>2026</copyright-year>
<copyright-holder>Ormaza-Gonz&#xe1;lez, Martillo-Bustamante, Espinoza, Proust, Michaud and Lebrun</copyright-holder>
<license>
<ali:license_ref start_date="2026-01-19">https://creativecommons.org/licenses/by/4.0/</ali:license_ref>
<license-p>This is an open-access article distributed under the terms of the <ext-link ext-link-type="uri" xlink:href="https://creativecommons.org/licenses/by/4.0/">Creative Commons Attribution License (CC BY)</ext-link>. The use, distribution or reproduction in other forums is permitted, provided the original author(s) and the copyright owner(s) are credited and that the original publication in this journal is cited, in accordance with accepted academic practice. No use, distribution or reproduction is permitted which does not comply with these terms.</license-p>
</license>
</permissions>
<abstract>
<p>Understanding the drivers of coastal ocean warming in the eastern Pacific is critical for distinguishing local variability from large&#x2011;scale climate phenomena such as El Ni&#xf1;o. This study tests the hypothesis that anomalous warming in northern Ecuadorian coastal waters during January 2024 was primarily driven by the seasonal intrusion of Panam&#xe1; Bay waters rather than the developing 2023&#x2013;2024 El Ni&#xf1;o event. Oceanographic and meteorological data were collected aboard the Pourquoi Pas? research vessel and compared against climatology (1940&#x2013;2024),  the 1997&#x2013;1998 El Ni&#xf1;o, and satellite observations. Winds were predominantly south-westerly, exceeding climatological averages while sea surface temperatures showed a pronounced south&#x2013;north gradient. Salinity and mixed&#x2011;layer depths remained within seasonal ranges. The Intertropical Convergence Zone persisted north. Comparative analysis revealed weak or absent El Ni&#xf1;o signals, with no discernible climatic impacts, wind, rain, thermocline and 20 C depths, T/S relationships, which were within seasonal values corroborated with biological and meteorological indicators. Instead, the seasonal incursion of Panam&#xe1; Bay waters was the dominant warming driver. These findings refine understanding of eastern Pacific variability and provide the first deep profiles from this region, including serendipitous evidence of Antarctic Intermediate Water at ~1000 m, thereby enhancing observational coverage in a historically undersampled area.</p>
</abstract>
<kwd-group>
<kwd>Eastern Pacific</kwd>
<kwd>El Ni&#xf1;o</kwd>
<kwd>halocline</kwd>
<kwd>Panam&#xe1; Bay</kwd>
<kwd>thermocline</kwd>
</kwd-group>
<funding-group>
<funding-statement>The author(s) declared that financial support was not received for this work and/or its publication.</funding-statement>
</funding-group>
<counts>
<fig-count count="16"/>
<table-count count="2"/>
<equation-count count="0"/>
<ref-count count="81"/>
<page-count count="20"/>
<word-count count="9568"/>
</counts>
<custom-meta-group>
<custom-meta>
<meta-name>section-at-acceptance</meta-name>
<meta-value>Coastal Ocean Processes</meta-value>
</custom-meta>
</custom-meta-group>
</article-meta>
</front>
<body>
<sec id="s1" sec-type="intro">
<label>1</label>
<title>Introduction</title>
<p>El Ni&#xf1;o is a coupled ocean-atmosphere phenomenon characterized by interannual variability and non-periodic occurrences (<xref ref-type="bibr" rid="B46">McPhaden et&#xa0;al., 2006</xref>; <xref ref-type="bibr" rid="B7">Cai et&#xa0;al., 2015</xref>; <xref ref-type="bibr" rid="B72">Wang et al., 2017</xref>; <xref ref-type="bibr" rid="B41">Li et&#xa0;al., 2024</xref>; <xref ref-type="bibr" rid="B33">Islam, 2025</xref>; <xref ref-type="bibr" rid="B71">Vialard et&#xa0;al., 2025</xref>; <xref ref-type="bibr" rid="B36">Kent et&#xa0;al., 2025</xref>; and references therein). It typically originates in the central equatorial Pacific, specifically within region 3.4 (bounded by 5&#xb0;N&#x2013;5&#xb0;S, 120&#xb0;W&#x2013;170&#xb0;W, see <xref ref-type="bibr" rid="B3">BoM, 2025</xref>), where heat accumulates in the water column (<xref ref-type="bibr" rid="B70">Trenberth, 1997</xref>; <xref ref-type="bibr" rid="B73">Wang and Fiedler, 2006</xref>; <xref ref-type="bibr" rid="B55">Ormaza-Gonz&#xe1;lez, 2016</xref>). This thermal anomaly is transported eastward by Kelvin waves, which elevate sea surface temperatures (SSTs) to 28&#x2013;29 &#xb0;C and deepen the thermocline to depths of approximately 150 meters (<xref ref-type="bibr" rid="B29">Halpern, 1987</xref>; <xref ref-type="bibr" rid="B26">Garc&#xe9;s-Vargas et&#xa0;al., 2005</xref>). The resulting thermal expansion rises mean sea level (MSL) by over 40 cm in region 1 + 2 (0-10S, 80W-90W).</p>
<p>The development of El Ni&#xf1;o significantly alters atmospheric dynamics. Westerly winds begin to dominate over the prevailing easterlies, and the redistribution of heat&#x2014;mediated by far-west Kelvin waves&#x2014;weakens the southern trade winds, thereby affecting the position of the Intertropical Convergence Zone (ITCZ). In regions 3 and 1 + 2 (<xref ref-type="bibr" rid="B3">BoM, 2025</xref>) the ITCZ shifts southward from its typical location near 5&#xb0;N to latitudes around 5&#x2013;10&#xb0;S. This displacement enhances evaporation and promotes persistent cloud formation, creating a greenhouse-like environment that reinforces heat accumulation. As a result, SSTs, evaporation rates, and rainfall intensify across the coastal zones of Ecuador and northern Peru (<xref ref-type="bibr" rid="B68">Thielen et&#xa0;al., 2023</xref>).</p>
<p>These meteorological and oceanographic ENSO events have profound socio-economic impacts at local, regional, and global scales (<xref ref-type="bibr" rid="B31">Huang et&#xa0;al., 2025</xref>). For example, during the 1997&#x2013;1998 El Ni&#xf1;o event, losses in Ecuador exceeded $2 billion USD, with nearly 300 fatalities, widespread destruction of infrastructure, landslides, urban flooding, and severe damage to agricultural lands (<xref ref-type="bibr" rid="B28">Glantz, 2001</xref>). Nearly 121 years before this event, another event was recorded in 1877&#x2013;1878, which caused enormous impact not only on the coasts of Ecuador and Per&#xfa; but also intense droughts in Asia, South-East Asia, and Africa triggering deaths over 20 million people (Aceituno et&#xa0;al., 2009). More recently <xref ref-type="bibr" rid="B31">Huang et&#xa0;al. (2025)</xref> have demonstrated how ENSO events cause quakes in the worldwide energy market with collateral impacts.</p>
<p>A more detailed consideration of the oceanographic dynamics in Bah&#xed;a de Panam&#xe1; is essential to contextualize the broader circulation patterns described in this study. The annual upwelling event in the Gulf of Panam&#xe1;, occurring between January and April, plays a critical role in modulating water masses and driving the transport of warmer waters toward the Equator. This process is closely linked to Atlantic tropical trade winds (<xref ref-type="bibr" rid="B54">Ordo&#x301;&#xf1;ez-Zu&#x301;&#xf1;iga et al., 2021</xref>), which blow east to west across Panam&#xe1; Bay and displace warm (27&#x2013;29 &#xb0;C) and relatively less saline (&lt;30&#x2013;32) surface waters (<xref ref-type="bibr" rid="B18">D&#x2019;Croz. et&#xa0;al., 1991</xref>). These waters are advected southward along the coasts of Colombia, Ecuador, and northern Peru, shaping regional circulation and productivity patterns (<xref ref-type="bibr" rid="B57">Ormaza-Gonz&#xe1;lezCede&#xf1;o, 2017</xref>; <xref ref-type="bibr" rid="B42">L&#xfc;bbecke et al., 2019</xref>).</p>
<p>Recent analyses confirm that the strength of these winds is fundamental to sustaining Panam&#xe1;&#x2019;s seasonal upwelling, with weakening events leading to suppression of nutrient&#x2212;rich waters and cascading ecological impacts (<xref ref-type="bibr" rid="B53">O&#x2019;Dea et&#xa0;al., 2025</xref>). Wind&#x2212;driven dynamics generate strong upwelling that brings nutrients (N, P, Si) from deeper layers to the surface, enriching the trophic chain and sustaining high biological productivity (<xref ref-type="bibr" rid="B18">D&#x2019;Croz. et&#xa0;al., 1991</xref>). Complementary studies emphasize the value of integrating remote sensing with <italic>in situ</italic> and biological data to capture these processes more effectively. For instance, satellite monitoring near Coiba Island enhances environmental and oceanographic assessments (<xref ref-type="bibr" rid="B16">Crawford et&#xa0;al., 2024</xref>), while meso&#x2212;zooplankton have been highlighted as biological indicators of climate and oceanographic variability in the Coiba Archipelago (<xref ref-type="bibr" rid="B5">Brugnoli et&#xa0;al., 2023</xref>).</p>
<p>Historical evidence underscores their importance of these warm and less saline waters southward intrusions. The 1925 Coastal El Ni&#xf1;o produced catastrophic flooding in northern Peru and Ecuador and is considered one of the strongest coastal warming events of the 20th century (<xref ref-type="bibr" rid="B66">Takahashi and Mart&#xed;nez, 2019</xref>). More recently, the 2017 Coastal El Ni&#xf1;o (January&#x2013;April) was driven by sudden Panam&#xe1; Bay intrusions, generating SST anomalies exceeding +2.6 &#xb0;C and torrential rainfall across Ecuador and Peru. In Peru, the event caused more than 100 fatalities, displaced over 300,000 people, health issues and inflicted economic losses surpassing $3 billion USD (<xref ref-type="bibr" rid="B20">Echevin et&#xa0;al., 2018</xref>; <xref ref-type="bibr" rid="B64">Rollenbeck et&#xa0;al., 2022</xref>; <xref ref-type="bibr" rid="B78">Yglesias-Gonz&#xe1;lez et&#xa0;al., 2023</xref>). Ecuador experienced extensive flooding in Guayas, Manab&#xed;, and Esmeraldas provinces, with severe agricultural losses in bananas, cacao, and rice, alongside widespread infrastructure damage. Importantly, <xref ref-type="bibr" rid="B57">Ormaza-Gonz&#xe1;lez and Cede&#xf1;o (2017)</xref> documented how Panam&#xe1; Bay intrusions during this period produced rainfall and warming patterns similar to El Ni&#xf1;o, reinforcing the need to consider these seasonal processes as independent drivers of coastal anomalies. These cases demonstrate that Panam&#xe1; Bay intrusions can generate impacts comparable to major ENSO events, yet remain underrepresented in global monitoring frameworks.</p>
<p>In 2022, Ormaza&#x2212;Gonz&#xe1;lez et&#xa0;al. predicted the development of an El Ni&#xf1;o event during 2023&#x2013;2024. By early 2023, a rare and intense SST anomaly was observed along the coasts of Ecuador and Peru (<xref ref-type="bibr" rid="B61">Peng et&#xa0;al., 2024</xref>), prompting global agencies to declare El Ni&#xf1;o conditions. In June 2023, NOAA confirmed El Ni&#xf1;o based on the Oceanic Ni&#xf1;o Index (ONI), with Kelvin waves propagating eastward and the Southern Oscillation Index (SOI) falling below -7. By September, both ONI and SOI exceeded thresholds and remained coupled for over five months, leading NOAA, BoM, and the UK Met Office to confirm the El Ni&#xf1;o 2023&#x2013;2024 event. By late December, the sixth consecutive warm Kelvin wave had reached region 1 + 2, accompanied by weakening trade winds and a southward&#x2212;shifting ITCZ.</p>
<p>In December 2023, El Ni&#xf1;o conditions prevailed, with ONI at 1.8 &#xb0;C and sea surface temperature anomalies (SSTA) reaching 2.0 &#xb0;C, classifying the event as strong. However, slight decreases in anomalies in the western equatorial Pacific were reported (<xref ref-type="bibr" rid="B48">MetOffice, 2024</xref>; <xref ref-type="bibr" rid="B52">NOAA, 2023</xref>), as cold waters began surfacing in the east. This apparent contradiction between global indices and local anomalies underscores the need to disentangle basin&#x2212;scale ENSO signals from regional processes such as Panam&#xe1; Bay intrusions &#x2014; the central motivation for this study.</p>
<p>While ENSO dynamics have been extensively documented, with abundant literature spanning physical mechanisms, teleconnections, and socio-economic impacts, references addressing seasonal Panam&#xe1; Bay intrusions remain comparatively scarce. Apart from a few regional studies (e.g., <xref ref-type="bibr" rid="B57">Ormaza-Gonz&#xe1;lez and Cede&#xf1;o, 2017</xref>; <xref ref-type="bibr" rid="B20">Echevin et&#xa0;al., 2018</xref>; <xref ref-type="bibr" rid="B67">Tan et&#xa0;al., 2024</xref>), the processes and impacts of Panam&#xe1; Bay waters are underrepresented in global climate analyses, despite their capacity to generate catastrophic events such as the 1925 and 2017 Coastal El Ni&#xf1;os, see <xref ref-type="bibr" rid="B27">Garreaud, (2018)</xref>. This imbalance in the scientific record underscores the need for focused research on Panam&#xe1; Bay intrusions, which can mimic or even amplify ENSO-like impacts but are often overlooked in monitoring frameworks.</p>
<p>The objective of this study is to disentangle the relative contributions of ENSO forcing and seasonal Panam&#xe1; Bay intrusions to the anomalous coastal warming observed in 2023&#x2013;2024. To achieve this, we present the first <italic>in situ</italic> oceanographic dataset of surface and vertical profiles extending to 2000 m depth along the easternmost equatorial Pacific margin, spanning the southern and northern coastal boundaries of Colombia and Ecuador. This dataset fills a critical observational gap in a historically under&#x2212;sampled region and provides the basis for testing our central hypothesis: that the 2023&#x2013;2024 coastal warming was primarily driven by Panam&#xe1; Bay intrusions rather than ENSO, and that El Ni&#xf1;o 2023&#x2013;2024 did not impact region 1 + 2 as strongly as the benchmark events of 1982&#x2013;1983 and 1997&#x2013;1998.</p>
<p>Although not the primary focus of this study, we also present and briefly analyze deep temperature profiles and temperature&#x2013;salinity (T/S) diagrams&#x2014;datasets rarely available for this region. These observations underscore the chronic lack of high&#x2212;resolution vertical oceanographic data in the eastern equatorial Pacific and provide critical context for disentangling El Ni&#xf1;o&#x2212;driven warming from seasonal Panam&#xe1; Bay influences. In doing so, they establish a valuable baseline for future climate modeling, predictive frameworks, and coastal risk assessments.</p>
</sec>
<sec id="s2" sec-type="materials|methods">
<label>2</label>
<title>Material and methods</title>
<sec id="s2_1">
<label>2.1</label>
<title><italic>In situ</italic> oceanographic and meteorological measurements</title>
<p>Surface Sea Temperature (SST), Salinity (S), Surface Air Temperature (SAT), wind speed and direction, atmospheric pressure (AP), and cloud cover were measured every two hours along the vessel&#x2019;s track from January 8 to 29, 2024, totaling &gt;250 <italic>in situ</italic> real time measurements (see <xref ref-type="fig" rid="f1"><bold>Figure 1</bold></xref>),. Cloud characteristics&#x2014;including type, height, and coverage&#x2014;were visually estimated following standard meteorological protocols. All measurements were conducted aboard the French research vessel Pourquoi pas? (<xref ref-type="bibr" rid="B12">Charcot, 1910</xref>), equipped with advanced oceanographic instrumentation (see vessel specifications at: <ext-link ext-link-type="uri" xlink:href="https://www.ifremer.fr/en/flotte-oceanographique-francaise/decouvrez-les-navires-de-la-flotte-oceanographique-francaise/le-pourquoi-pas">https://www.ifremer.fr/en/flotte-oceanographique-francaise/decouvrez-les-navires-de-la-flotte-oceanographique-francaise/le-pourquoi-pas</ext-link>). The position of the Intertropical Convergence Zone (ITCZ) was monitored using freely available meteorological platforms, including <ext-link ext-link-type="uri" xlink:href="http://wxcharts.com">wxcharts.com</ext-link>, which provided real-time synoptic maps and atmospheric overlays.</p>
</sec>
<sec id="s2_2">
<label>2.2</label>
<title>Vertical profiling and instrumentation</title>
<p>Vertical Sea Temperature (VST) profiles were obtained using Expendable Bathythermographs (XBTs; Sippican Deep Blue series, Lockheed Martin), with a precision of &#xb1;0.2 &#xb0;C (<xref ref-type="bibr" rid="B32">IOC, 2018</xref>; <xref ref-type="bibr" rid="B59">Parks et&#xa0;al., 2022</xref>). Multiple XBT models were deployed depending on the vessel&#x2019;s speed and target depth (0&#x2013;2000 m); during this cruise, the R/V Pourquoi pas? maintained a maximum speed of 5 knots. A total of 37 XBT casts were conducted at designated stations (see <xref ref-type="fig" rid="f1"><bold>Figure&#xa0;1</bold></xref>), spanning the northern Ecuadorian and southern Colombian coastal zones.</p>
<fig id="f1" position="float">
<label>Figure&#xa0;1</label>
<caption>
<p>Tracks and the positions sites of continuous surface sampling (blue dots) and XBT launches (red stars) from the 8th to 29<sup>th</sup> of January. The sequential sampling was from south to north. Typical ship speed: 5 knots (2.57 m/s). Source: Authors' own elaboration. Based on unpublished preliminary report; see also <xref ref-type="bibr" rid="B50">Michaud et al. (2024)</xref>.</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g001.tif">
<alt-text content-type="machine-generated">Map showing oceanographic data collection points along the Pacific coast of Ecuador. Red stars indicate XBT_SIPPICAN locations, and blue circles represent temperature and salinity measurement sites. Black lines trace the navigation route. Inset map highlights the region within South America's northwest, near Quito, Ecuador. The legend and scale are included.</alt-text>
</graphic></fig>
<p>Geolocation and surface meteorological data were acquired from onboard instruments integrated into the vessel&#x2019;s navigation and data acquisition systems. These included GPS, thermosalinographs, anemometers, barometers, and cloud observation protocols consistent with World Meteorological Organization standards.</p>
</sec>
<sec id="s2_3">
<label>2.3</label>
<title>Satellite and reanalysis data</title>
<p>To complement <italic>in situ</italic> observations, satellite-derived data from NASA&#x2019;s Terra (EOS AM-1) and Aqua (EOS PM-1) platforms were used. These satellites carry the Moderate Resolution Imaging Spectroradiometer (MODIS), which provided temporal and spatial measurements of chlorophyll-a concentrations and the diffuse attenuation coefficient (Kd at 490 nm) during the study period. Additionally, reanalysis products were accessed via the Copernicus Marine Environment Monitoring Service (CMEMS), specifically the WIND_GLO_PHY_CLIMATE_L4_MY_012_003 dataset. This product includes climatological wind fields from January 1993 to 2025, enabling historical comparisons and anomaly detection.</p>
</sec>
<sec id="s2_4">
<label>2.4</label>
<title>Supplementary observations and navigation tracks</title>
<p>Supplementary regional meteorological and oceanographic insights were gathered from verified social media sources, including real-time observations shared by local agencies and research institutions during the cruise period. These informal data points were used to cross-reference <italic>in situ</italic> and satellite observations.</p>
<p>The Pourquoi pas? followed a predefined navigation track designed for seismic, bathymetric, and oceanographic profiling. These tracks closely mirrored those of previous cruises in the region (e.g., <xref ref-type="bibr" rid="B49">Michaud et&#xa0;al., 2015</xref>), ensuring continuity in spatial coverage and facilitating comparative analysis.</p>
</sec>
<sec id="s2_5">
<label>2.5</label>
<title>Data and graphic analysis</title>
<p>To complement the <italic>in situ</italic> measurements collected during the cruise, third-party datasets were incorporated to enhance spatial and temporal coverage. These included satellite observations, reanalysis products, and publicly available meteorological data (see <xref ref-type="table" rid="T1"><bold>Table&#xa0;1</bold></xref> and references).</p>
<table-wrap id="T1" position="float">
<label>Table&#xa0;1</label>
<caption>
<p>Summary of spatially interpolated variables. See links in reference section.</p>
</caption>
<table frame="hsides">
<thead>
<tr>
<th valign="middle" align="center">References (links)</th>
<th valign="middle" align="center">Variables</th>
<th valign="middle" align="center">Sources</th>
</tr>
</thead>
<tbody>
<tr>
<td valign="middle" align="center"><ext-link ext-link-type="uri" xlink:href="https://data.marine.copernicus.eu/product/GLOBAL_MULTIYEAR_PHY_001_030/description">1</ext-link></td>
<td valign="middle" align="center">Sea Surface Temperature</td>
<td valign="middle" align="center">CMEMS - MERCATOR GLORYS12V1</td>
</tr>
<tr>
<td valign="middle" align="center"><ext-link ext-link-type="uri" xlink:href="https://data.marine.copernicus.eu/product/GLOBAL_MULTIYEAR_PHY_001_030/description">2</ext-link></td>
<td valign="middle" align="center">Sea Surface Salinity</td>
<td valign="middle" align="center">CMEMS - MERCATOR GLORYS12V1</td>
</tr>
<tr>
<td valign="middle" align="center"><ext-link ext-link-type="uri" xlink:href="https://data.marine.copernicus.eu/es/product/GLOBAL_MULTIYEAR_BGC_001_029/description">3</ext-link></td>
<td valign="middle" align="center">Chlorophyll - a</td>
<td valign="middle" align="center">CMEMS - Global Ocean Biogeochemistry Hindcast</td>
</tr>
<tr>
<td valign="middle" align="center"><ext-link ext-link-type="uri" xlink:href="https://cds.climate.copernicus.eu/datasets/reanalysis-era5-single-levels-monthly-means?tab=overview">4</ext-link></td>
<td valign="middle" align="center">Air Temperature [2m]</td>
<td valign="middle" align="center">ECMWF - ERA5</td>
</tr>
<tr>
<td valign="middle" align="center"><ext-link ext-link-type="uri" xlink:href="https://cds.climate.copernicus.eu/datasets/reanalysis-era5-single-levels-monthly-means?tab=overview">5</ext-link></td>
<td valign="middle" align="center">Atmospheric Pressure (Surface)</td>
<td valign="middle" align="center">ECMWF - ERA5</td>
</tr>
<tr>
<td valign="middle" align="center"><ext-link ext-link-type="uri" xlink:href="https://data.marine.copernicus.eu/product/WIND_GLO_PHY_CLIMATE_L4_MY_012_003/description">6</ext-link></td>
<td valign="middle" align="center">Wind Speed</td>
<td valign="middle" align="center">CMEMS - Global Ocean Monthly Mean Sea Surface Wind and Stress from Scatterometer and Model</td>
</tr>
<tr>
<td valign="middle" align="center"><ext-link ext-link-type="uri" xlink:href="https://data.marine.copernicus.eu/product/MULTIOBS_GLO_PHY_TSUV_3D_MYNRT_015_012/description">7</ext-link></td>
<td valign="middle" align="center">Mixed Layer Depth</td>
<td valign="middle" align="center">CMEMS - Multi Observation Global Ocean ARMOR3D</td>
</tr>
<tr>
<td valign="middle" align="center"><ext-link ext-link-type="uri" xlink:href="https://cds.climate.copernicus.eu/datasets/reanalysis-oras5?tab=overview">8</ext-link></td>
<td valign="middle" align="center">Isotherm 20 &#xb0;C Depth</td>
<td valign="middle" align="center">ECMWF &#x2013; ORAS5</td>
</tr>
</tbody>
</table>
<table-wrap-foot>
<fn>
<p><sup>[1, 2]</sup><ext-link ext-link-type="uri" xlink:href="https://data.marine.copernicus.eu/product/GLOBAL_MULTIYEAR_PHY_001_030/description">https://data.marine.copernicus.eu/product/GLOBAL_MULTIYEAR_PHY_001_030/description</ext-link></p></fn>
<fn>
<p><sup>[3]</sup><ext-link ext-link-type="uri" xlink:href="https://data.marine.copernicus.eu/es/product/GLOBAL_MULTIYEAR_BGC_001_029/description">https://data.marine.copernicus.eu/es/product/GLOBAL_MULTIYEAR_BGC_001_029/description</ext-link></p></fn>
<fn>
<p><sup>[4, 5]</sup><ext-link ext-link-type="uri" xlink:href="https://cds.climate.copernicus.eu/datasets/reanalysis-era5-single-levels-monthly-means?tab=overview">https://cds.climate.copernicus.eu/datasets/reanalysis-era5-single-levels-monthly-means?tab=overview</ext-link></p></fn>
<fn>
<p><sup>[6]</sup><ext-link ext-link-type="uri" xlink:href="https://data.marine.copernicus.eu/product/WIND_GLO_PHY_CLIMATE_L4_MY_012_003/description">https://data.marine.copernicus.eu/product/WIND_GLO_PHY_CLIMATE_L4_MY_012_003/description</ext-link></p></fn>
<fn>
<p><sup>[7]</sup><ext-link ext-link-type="uri" xlink:href="https://data.marine.copernicus.eu/product/MULTIOBS_GLO_PHY_TSUV_3D_MYNRT_015_012/description">https://data.marine.copernicus.eu/product/MULTIOBS_GLO_PHY_TSUV_3D_MYNRT_015_012/description</ext-link></p></fn>
<fn>
<p><sup>[8]</sup><ext-link ext-link-type="uri" xlink:href="https://cds.climate.copernicus.eu/datasets/reanalysis-oras5?tab=overview">https://cds.climate.copernicus.eu/datasets/reanalysis-oras5?tab=overview</ext-link></p></fn>
</table-wrap-foot>
</table-wrap>
<p>Spatial interpolation of oceanographic variables was performed using the Kriging method, a geostatistical technique widely applied in earth and environmental sciences. Kriging is particularly effective for interpolating irregularly distributed data when initial estimates or known autocorrelation structures are unavailable (<xref ref-type="bibr" rid="B30">Hansen and Poulain, 1995</xref>; <xref ref-type="bibr" rid="B37">Kusuma et&#xa0;al., 2018</xref>). The method leverages spatial dependence among observations to estimate values at unsampled locations, making it ideal for modeling continuous environmental fields from discrete point measurements (<xref ref-type="bibr" rid="B38">Le and Zidek, 2006</xref>).</p>
<p>Each variable&#x2014;such as sea surface temperature, salinity, and wind speed&#x2014;was sourced from distinct datasets, including <italic>in situ</italic> field measurements and satellite-derived or reanalysis products. These variables were individually interpolated using Kriging to generate continuous spatial maps suitable for comparative analysis across the study region. A summary of the interpolated variables and their sources is provided in <xref ref-type="table" rid="T1"><bold>Table&#xa0;1</bold></xref>.</p>
</sec>
<sec id="s2_6">
<label>2.6</label>
<title>Research area</title>
<p>The study area (see map) extends from 0&#xb0; 0.24&#x2032; S to 2&#xb0; 02&#x2032; N and from 79&#xb0; 19&#x2032; W to 81&#xb0; 12&#x2032; W, encompassing the southern coast of Colombia and the northern coast of Ecuador. This region is characterized by pronounced oceanographic and meteorological dynamics, and is also notable for its seismological activity and sediment biogeochemical processes.</p>
</sec>
<sec id="s2_7">
<label>2.7</label>
<title>Oceanographic and climatic dynamics</title>
<p>Surface ocean and coastal circulation in this region is primarily influenced by the Panam&#xe1; Bight current system (0&#x2013;9&#xb0; N; 73&#x2013;90&#xb0; W), which comprises three major currents with typical velocities ranging from 0.1 to 0.5 m s<sup>-1</sup> (see: earth:: a global map of wind, weather, and ocean conditions). These include:</p>
<list list-type="bullet">
<list-item>
<p>The eastward-flowing North Equatorial Counter Current (NECC),</p></list-item>
<list-item>
<p>The near-circular Panam&#xe1; Bight Cyclonic Gyre (PBCG),</p></list-item>
<list-item>
<p>The westward-flowing South Equatorial Current (SEC) (<xref ref-type="bibr" rid="B9">Chaigneau et&#xa0;al., 2006</xref>).</p></list-item>
</list>
<p>The west Pacific North Equatorial Countercurrent (NECC) has been shown to dramatically influence the impact of El Ni&#xf1;o events (1987-1988) on the western coasts of Colombia, Ecuador, and Per&#xfa; primarily. During 1982-1983, 1987-1988, 2009&#x2013;2010 El Ni&#xf1;o events, the NECC jet intensity proved to be decisive on the coastal impact (<xref ref-type="bibr" rid="B76">Wyrtki, 1973</xref>, <xref ref-type="bibr" rid="B77">Wyrtki, 1974</xref>; <xref ref-type="bibr" rid="B81">Zhao et&#xa0;al., 2013</xref>; <xref ref-type="bibr" rid="B75">Webb, 2018</xref>).</p>
<p>The South Equatorial Current (SEC) is a major westward-flowing surface current in the tropical Pacific Ocean, driven primarily by the trade winds. It transports warm, nutrient-poor waters across the equatorial region and plays a key role in upper ocean circulation and heat distribution. According to <xref ref-type="bibr" rid="B9">Chaigneau et&#xa0;al. (2006)</xref>, the SEC exhibits complex spatial variability influenced by mesoscale structures and interactions with equatorial dynamics.</p>
<p>The PBCG&#x2014;often referred to as the Panam&#xe1; Bight Current&#x2014;transports warm (&gt;27 &#xb0;C) and relatively low-salinity (32&#x2013;33) surface waters to the northern coast of Ecuador (around 2&#xb0; N). This influx triggers rapid coastal warming that can extend southward to southern Ecuador and northern Peru (3&#x2013;5&#xb0; S), as noted by <xref ref-type="bibr" rid="B57">Ormaza-Gonz&#xe1;lez and Cede&#xf1;o (2017)</xref>. This phenomenon is also known in Per&#xfa; as El Ni&#xf1;o Costero (<xref ref-type="bibr" rid="B63">Ram&#xed;rez and Briones, 2017</xref>; <xref ref-type="bibr" rid="B23">Espinoza-Morriber&#xf3;n et&#xa0;al., 2021</xref>) and typically occurs seasonally from January to April&#x2013;May, depending on the strength of the Panam&#xe1; Bight Current. Its impacts resemble those of the broader El Ni&#xf1;o event.</p>
<p>Conversely, the northward-flowing Humboldt Current (<xref ref-type="bibr" rid="B10">Chaigneau et&#xa0;al., 2013</xref>) can extend beyond the equator, carrying cold waters (typically &lt;19 &#xb0;C) with salinity levels above 35. It predominates along the Ecuadorian coast from May to December. The interplay between the PBCG and the Humboldt Current defines two distinct seasons: a rainy season from January to April and a dry season from May to December, with precipitation levels significantly above or below average, respectively. These seasonal dynamics also affect marine productivity&#x2014;diminished during the rainy season and enhanced during the dry season. When El Ni&#xf1;o events reach the coasts of Ecuador and Peru, they often result in catastrophic, flood-inducing rainfall. Understanding the interplay between these oceanographic drivers is essential for distinguishing seasonal warming from El Ni&#xf1;o-related anomalies&#x2014;an objective central to this study.</p>
<p>El Ni&#xf1;o is officially declared when Sea Surface Temperature Anomalies (SSTA) in region 3.4 (5&#xb0; N&#x2013;5&#xb0; S, 120&#x2013;170&#xb0; W) exceed +0.5 &#xb0;C for at least five consecutive months, accompanied by a Southern Oscillation Index (SOI) below &#x2212;7 (<xref ref-type="bibr" rid="B1">Adamson, 2022</xref>). The accumulated thermal energy in this region is gradually transmitted eastward over 2&#x2013;3 months via descending warm Kelvin waves, which elevate sea surface temperatures along the Ecuadorian coast. This is an interannual phenomenon.</p>
</sec>
</sec>
<sec id="s3" sec-type="results">
<label>3</label>
<title>Results</title>
<sec id="s3_1">
<label>3.1</label>
<title>Meteorology</title>
<sec id="s3_1_1">
<label>3.1.1</label>
<title>Surface air temperature</title>
<p><italic>In situ</italic> SAT fluctuated around 26.5 &#xb1; 0.87 C ranging from 24.3C to 28.9C from southern to northern latitudes respectively. The distribution clearly shows an increasing SAT from south to north, the tongue-like distribution shows colder surface water intruding north coast as trade winds were blowing consistently from the south. Similar SAT pattern for same month 2024 is observed from data reported from 1940 to 2024, however the climatology shows colder SAT as it ranged from 25.9 C to 26.7 C, with a media value of 26.34C. The red dots at the same geographical position provided punctual STA, thus the most northern-southern values are 27.1 - 25.3 and 25.5 &#x2013; 24.5 C for <italic>in situ</italic> and climatology data, respectively.</p>
<p>During January 2024, atmospheric pressure ranged from 1002.8 to 1011.1 hPa, showing a variation over 8.3 hPa with an average of 1007.3 &#xb1; 1.77 hPa. The lowest values were recorded in the north and the highest in the south. In contrast, the January climatological average for 1940&#x2013;2022 exhibited slightly higher values, ranging from 1008.9 hPa in the south to 1011.3 hPa in the north; i.e, a difference of 2.4 hPa. Illustrative values (red dots) show <italic>in situ</italic> and climatological pressures of 1005.1 and 1010.8 hPa in the north, and 1008.2 and 1008.9 hPa in the south, respectively. Overall, <italic>in situ</italic> atmospheric pressure in 2024 was lower than the climatological average, though it followed the same spatial distribution pattern.</p>
</sec>
<sec id="s3_1_2">
<label>3.1.2</label>
<title>Surface winds</title>
<p>Wind speeds ranged from 0.41 to 11.47 m/s (0.8 to 22.3 knots), with an average of 5.6 &#xb1; 2.0 m/s. The lowest values were recorded in the southern coastal areas, while the highest occurred over the open ocean between 1&#xb0;S and 1&#xb0;N, west of 81&#xb0;W. Overall, the strongest winds were observed north of Ecuador. Wind direction predominantly ranged from 200&#xb0; to 320&#xb0;, with a median value of 225&#xb0; (<xref ref-type="fig" rid="f2"><bold>Figure&#xa0;2</bold></xref>). In contrast, satellite climatological data from 1995 to 2024 reported a median wind speed of 3.61 &#xb1; 0.52 m/s (2.33&#x2013;4.28 m/s), which is typical for this time of year, and showed a similar directional pattern. Notably, there is a marked difference of nearly 2.00 m/s between the <italic>in situ</italic> measurements and the climatological averages.</p>
<fig id="f2" position="float">
<label>Figure&#xa0;2</label>
<caption>
<p>Wind speed and direction. Left. <italic>In situ</italic>, January 2024. Right. Climatology 1995-2024. Direction in degrees (arrows).</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g002.tif">
<alt-text content-type="machine-generated">Side-by-side maps about surface winds (speed and direction) on the eastern Pacific Ocean. The left map shows in-situ data with directions depicted by arrows. The right map shows climatology data, similarly labeled. Color gradients indicate current strength from blue (weak) to red (strong). Both maps include latitude and longitude markers, a scale bar, and a compass for orientation.</alt-text>
</graphic></fig>
</sec>
<sec id="s3_1_3">
<label>3.1.3</label>
<title>The Inter Tropical Convergence Zone and cloudiness</title>
<p>On average, throughout the cruise, skies were predominantly cloudy (6/8 coverage), with medium-altitude clouds (approximately 2000 m) consisting mainly of cumulus and fracto-stratus formations. On some days, cloud cover intensified into nimbostratus (see photos in <xref ref-type="fig" rid="f3"><bold>Figure&#xa0;3</bold></xref>). Meanwhile, the most eastern southern flank of the Intertropical Convergence Zone (ITCZ) was monitored using freely available meteorological platforms such as <ext-link ext-link-type="uri" xlink:href="http://wxcharts.com">wxcharts.com</ext-link>. It was generally located between 2&#xb0;N and 3&#xb0;N, occasionally shifting southward to nearly 1&#xb0;N (see <ext-link ext-link-type="uri" xlink:href="https://www.ventusky.com/?p=3.7;-85.3;4&amp;l=rain-3h&amp;t=20240113/1200">https://www.ventusky.com/?p=3.7;-85.3;4&amp;l=rain-3h&amp;t=20240113/1200</ext-link>, 10 January 2024).</p>
<fig id="f3" position="float">
<label>Figure&#xa0;3</label>
<caption>
<p>Typical cloud cast from the 8th to the 29th of January. <bold>(A, B)</bold> cumulus stratus; <bold>(C, D)</bold> fracto stratus medium altitude (around 2000 m height). Credit: Authors.</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g003.tif">
<alt-text content-type="machine-generated">Four images showing different sea and sky views from a ship's deck labeled A, B, C, and D. A shows a calm sea under a cloudy sky with a hint of sunlight. B features the ship's bow pointing towards a vast ocean under a similar cloudy sky. C presents darker clouds with visible sun rays, and the ship's bow prominently in view. D displays a darker, overcast sky with a slightly rough sea.</alt-text>
</graphic></fig>
</sec>
</sec>
<sec id="s3_2">
<label>3.2</label>
<title>Oceanography. <italic>In situ</italic> measurements</title>
<sec id="s3_2_1">
<label>3.2.1</label>
<title>Sea surface temperature</title>
<p>Sea surface temperature (SST) ranged from 25.90 &#xb0;C in the southernmost part of the study area to 28.60 &#xb0;C in the north, revealing a distinct latitudinal gradient. A slight cooling trend was also evident from the coastal zone toward the open ocean (<xref ref-type="fig" rid="f4"><bold>Figure&#xa0;4</bold></xref>, left). Isotherms between 27 &#xb0;C and 28 &#xb0;C were aligned parallel to the coastline, extending from south to north, with the 28 &#xb0;C isotherms concentrated north of approximately 1.7&#xb0;N. The lowest SST values were recorded near 1&#xb0;S, between 81&#xb0;W and 82&#xb0;W.</p>
<fig id="f4" position="float">
<label>Figure&#xa0;4</label>
<caption>
<p>Sea Surface Temperature (SST). Left. <italic>In situ</italic> data taken. Right Climatology (1993-2021), from Copernicus. Red dots are at the same geographical position in both graphs.</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g004.tif">
<alt-text content-type="machine-generated">Two panels compare ocean temperatures. The left panel displays in-situ sea temperatures ranging from 25.9 to 28.4 degrees Celsius. The right panel shows climatology with temperatures from 26 to 26.6 degrees Celsius. Both maps indicate locations with red dots and have a color gradient from blue to yellow, representing cooler to warmer temperatures. Latitude and longitude markings are present, with a color scale indicating temperature in Celsius on the right. A compass rose in each panel indicates the north direction.</alt-text>
</graphic></fig>
<p>Climatological SST data for January (<xref ref-type="fig" rid="f4"><bold>Figure&#xa0;4</bold></xref>, right) showed the highest temperatures in the northeastern sector, around 26.5&#xb0;C, gradually decreasing to approximately 26.0&#xb0;C in the southern region. Notably, the isotherm distribution differed between the climatological averages and the <italic>in situ</italic> measurements from January 2024, with significantly warmer SSTs observed during the latter. For instance, SSTs reached 28.4 &#xb0;C compared to 26.5 &#xb0;C at the same location (see red dots), and 26.8 &#xb0;C versus 26.0 &#xb0;C in the southern portion.</p>
</sec>
<sec id="s3_2_2">
<label>3.2.2</label>
<title>Sea Surface Salinity</title>
<p>Sea Surface Salinity (SSS) ranged from 29.60 to 32.79, gradually increasing from north to south, with an average value of 31.60 &#xb1; 0.78 (<xref ref-type="fig" rid="f5"><bold>Figure&#xa0;5</bold></xref>, left). A localized low-salinity zone (~30.12) was identified near 2&#xb0;N, where a tongue of fresher water extended southward, effectively separating coastal (eastern) from offshore (western) waters. Salinity increased westward across this boundary.</p>
<fig id="f5" position="float">
<label>Figure&#xa0;5</label>
<caption>
<p>Sea surface salinity distribution in the researched area. Left. Data taken <italic>in situ</italic>. Right. climatology (1994-2021) from Copernicus data.</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g005.tif">
<alt-text content-type="machine-generated">Side-by-side maps titled &#x201c;In-Situ&#x201d; and &#x201c;Climatology&#x201d; show temperature variations off a coastline with latitude and longitude coordinates. Both maps display colored gradients from blue to orange, indicating temperatures from 28 to 34 degrees Celsius. Red dots with temperature values are plotted along the coast. The maps include a scale bar for distance.</alt-text>
</graphic></fig>
<p>Climatological SSS data (1991&#x2013;1994) from Copernicus revealed a broadly similar spatial pattern, with salinities (see red dots in <xref ref-type="fig" rid="f5"><bold>Figure&#xa0;5</bold></xref>) ranging from approximately 32.10 in the north to a maximum of 33.00 in the south. Compared to <italic>in situ</italic> measurements, the northern values were over 2 units higher (<italic>in situ</italic> ~30.00), while southern values differed by less than 0.71. This indicates a pronounced salinity gradient from south to north.</p>
</sec>
<sec id="s3_2_3">
<label>3.2.3</label>
<title>Deep vertical distribution of temperature and salinity</title>
<p>This study presents the first reported vertical profiles of temperature and salinity in the eastern equatorial Pacific, based on 37 XBT casts (see <xref ref-type="fig" rid="f1"><bold>Figure&#xa0;1</bold></xref> and SM1 for geographical positions). All profiles exhibited the classic vertical structure characteristic of tropical oceanic waters. The Pp5 profile (see <xref ref-type="fig" rid="f1"><bold>Figure&#xa0;1</bold></xref>) at the southern latitude, illustrates a representative distribution down to 100 m depth. The mixed layer, defined following <xref ref-type="bibr" rid="B34">Johnson and Lyman (2022)</xref>, extended to approximately 10 m. A pronounced thermocline was observed between 16 m and 41 m, with a temperature gradient of 0.33&#xb0;C/m, followed by a more gradual decline to 100 m. Salinity within the mixed layer varied slightly, ranging from 33.64 to 34.03.</p>
<p>In general, below the surface layer, temperature decreased progressively: reaching ~15&#xb0;C at 200 m, ~10&#xb0;C between 400&#x2013;500 m, and ultimately 2.29 &#xb0;C at 2000 m (<xref ref-type="fig" rid="f6"><bold>Figure&#xa0;6</bold></xref>). In contrast, salinity remained relatively constant at 35.00 from 50 m downward, however slightly lower salinity below 1000 m was found in deep profiles down to 2000 meters. These profiles represent the deepest temperature measurements reported for this region.</p>
<fig id="f6" position="float">
<label>Figure&#xa0;6</label>
<caption>
<p>Deep vertical profile pP28 at: 1 0&#x2019;N, 80 21&#x2019;W, (see SM1). 20/01/2024.</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g006.tif">
<alt-text content-type="machine-generated">Graph showing ocean depth versus temperature and salinity. The red line represents temperature in degrees Celsius, starting high at the surface and decreasing with depth. The blue dashed line represents salinity in practical salinity units (PSU), remaining relatively constant with slight variation at different depths. Depth is measured in meters along the vertical axis.</alt-text>
</graphic></fig>
</sec>
<sec id="s3_2_4">
<label>3.2.4</label>
<title>Deep latitudinal profiles of thermal structure</title>
<p><xref ref-type="fig" rid="f7"><bold>Figure&#xa0;7</bold></xref> presents four latitudinal transects derived from XBT data, illustrating the vertical distribution of the 28&#x2013;27 &#xb0;C isotherms. These isotherms reach their greatest depth in the northernmost profile (A), descending to approximately 30&#x2013;35 m, while in the southern profile they shoal to less than 12 m. Along transect B, the isotherms rise to around 20 m before deepening again to nearly 30 m in profile C, indicating pronounced spatial variability in thermal stratification.</p>
<fig id="f7" position="float">
<label>Figure&#xa0;7</label>
<caption>
<p>Latitudinal vertical profiles (1-100m). A: 1.49N-1.99N; B: 0.91N-1.16N; C: 0.01S &#x2013; 0.40N; and D: 0.89S &#x2013; 0.28S. The XBT launch codes are at the top of each graph.</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g007.tif">
<alt-text content-type="machine-generated">Four contour plots (A, B, C, D) depict temperature variations with depth and distance in a specific region. Each plot shows depth in meters on the vertical axis and distance in kilometers on the horizontal axis. Temperature ranges from 0.0&#xb0;C to 32.0&#xb0;C, indicated by a color gradient from green to red. Contour lines and labels within the plots specify temperature values at different depths and distances.</alt-text>
</graphic></fig>
<p>The steep thermal gradients observed in profile D suggest active vertical mixing, likely driven by turbulent processes. In contrast, profile C&#x2014;located near the equatorial line&#x2014;exhibits signs of downward displacement of surface waters, resulting in a deepened mixed layer. This feature coincides with reduced wind intensity at this latitude, which may have facilitated the southward intrusion of warm waters from Panama Bay. Profile B, meanwhile, appears to reflect an upward displacement of subsurface waters, contributing to a shallower mixed layer and the emergence of 20&#xb0;C waters at relatively shallow depths.</p>
<p>Further north, profile A again shows evidence of Panama Bay water intrusion, like the pattern observed in profile C. It is important to note that the whole dataset was not collected in real time; there is a temporal gap of approximately 20 days between the southernmost and northernmost profiles, which may influence the observed variability.</p>
<p><xref ref-type="fig" rid="f8"><bold>Figure&#xa0;8</bold></xref> shows longitudinal vertical profiles along three latitudes from 0 to 1000 meters. Overall, the isotherms show highly stratified water columns below 50&#x2013;70 meters depth, although there is a slight vertical displacement between isotherms 5&#x2013;10 C indicating presence of distinct water masses, probably the Antarctic Intermediate Water (AAIW), specifically the South Pacific subtype (<xref ref-type="bibr" rid="B4">Bostock et&#xa0;al., 2013</xref>). It is well known the AAIW is part of the global thermohaline circulation, which moves water masses across vast distances. The AAIW is advected northward via the Peru-Chile Undercurrent and the Equatorial Undercurrent system. Also, the complex bathymetry of the eastern Pacific, including ridges and continental slopes, can steer AAIW northward.</p>
<fig id="f8" position="float">
<label>Figure&#xa0;8</label>
<caption>
<p>Longitudinal vertical profiles along three latitudes from 0 to 1000 meters. The XBT code is at the top. See <xref ref-type="fig" rid="f1"><bold>Figure&#xa0;1</bold></xref> and SM1 for geographical position.</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g008.tif">
<alt-text content-type="machine-generated">Temperature profiles at three ocean transects (A, B, C) are shown with depth versus distance in kilometers. Color gradient from red to blue indicates temperatures from 32.0&#xb0;C to 0.0&#xb0;C. Each transect has labeled isotherms marking specific temperature depths.</alt-text>
</graphic></fig>
</sec>
<sec id="s3_2_5">
<label>3.2.5</label>
<title>Mixed layer and the vertical 20 &#xb0;C isotherm</title>
<p>During January 2024, the horizontal distribution of the mixed layer depth (MLD) and the 20 &#xb0;C isotherm (<xref ref-type="fig" rid="f9"><bold>Figure&#xa0;9</bold></xref>) reveals distinct spatial variability across the study area. The MLD averaged 22.4 m, ranging from 6.1 m to 40.7 m. Shallowest thermocline was between 1S and just below the equatorial line; a second shallow core was at 1N. The deepest mixed layers or thermoclines were observed near the northern part of the equatorial line and the coastal region (approximately 0.0 to 0.5&#xb0;N), suggesting enhanced vertical mixing or wind-driven processes in these areas, where the highest wind speed was measured. The 20&#xb0;C isotherm was located at an average depth of 37.9 m, spanning from 10.1 to 56.6 meters depth. Concurrently, the core of the deepest 20&#xb0;C isotherm coincided with the corresponding deeper thermocline, similar pattern occurred with shallow thermocline and 20 C isotherms. The spatial distribution of gradients suggests surface advection processes from north to south and vice versa on a wave like movement.</p>
<fig id="f9" position="float">
<label>Figure&#xa0;9</label>
<caption>
<p><italic>In situ</italic> mixed layer depth (left), and 20C isotherm depth (right) during January 2024.</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g009.tif">
<alt-text content-type="machine-generated">Two contour maps side by side show varying depths in meters, indicated by color gradients. The left map ranges from blue to green representing shallower depths. The right map ranges from green to red representing deeper depths. Both maps cover a similar geographic area, with latitude and longitude markers, and include a scale bar and compass directions.</alt-text>
</graphic></fig>
</sec>
<sec id="s3_2_6">
<label>3.2.6</label>
<title>Water masses</title>
<p>Temperature-Salinity (T-S) diagrams revealed the presence of distinct water masses during the sampling period. As shown in <xref ref-type="fig" rid="f10"><bold>Figure&#xa0;10</bold></xref>, three principal water masses were identified at the northernmost sampling site: Panam&#xe1; Bay Water, Humboldt Current Water, and Deep Pacific Water (see e.g., <xref ref-type="bibr" rid="B62">Peters et al., 2018</xref>). Among these, Panam&#xe1; Bay Water was the most prominent, occupying the surface layer and extending several meters downward throughout the study area. This water mass is characterized by elevated temperatures and reduced salinity, consistent with its coastal origin and influence from tropical surface waters.</p>
<fig id="f10" position="float">
<label>Figure&#xa0;10</label>
<caption>
<p>T-S diagram at the most northern sampling site. (Pp33, see Figure1, MS1 geographical position). Ellipses: Yellow, near-circular Panam&#xe1; Bight Cyclonic Gyre (PBCG) waters. Green, Humboldt current waters. Blue, deep ocean waters (Including AAIW). See SM2 for all XBT profiles.</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g010.tif">
<alt-text content-type="machine-generated">Graph depicting temperature versus salinity with depth as a color gradient. Data points are connected, forming a path across various temperature and salinity levels, annotated with dashed contours showing density lines. The path is highlighted with orange and green dashed lines at different segments, and a color bar below indicates depth ranging from zero to two thousand meters.</alt-text>
</graphic></fig>
<p>In contrast, Humboldt Current Water and Deep Pacific Water were confined to greater depths and exhibited more localized distributions, suggesting limited vertical mixing and a stratified water column during the observation period. At approximately 1000 m depth, the T-S correlation indicated a density exceeding 1.027 g/cm&#xb3;, consistent with the presence of Antarctic Intermediate Water (AAIW), specifically the South Pacific subtype (<xref ref-type="bibr" rid="B4">Bostock et&#xa0;al., 2013</xref>). Above this layer, a water mass with salinity near 35 and temperatures around 15 &#xb0;C was observed, likely associated with the westward-flowing South Equatorial Current (SEC) and the subsurface extension of Humboldt Current waters. As the characterization of water masses was not a primary objective of this study, the full set of temperature-salinity profiles (32 in total) is provided in the <xref ref-type="supplementary-material" rid="SM1"><bold>Supplementary Material</bold></xref>. This dataset constitutes the first reported T-S observations for this region of the eastern tropical Pacific.</p>
</sec>
</sec>
</sec>
<sec id="s4" sec-type="discussion">
<label>4</label>
<title>Discussion</title>
<p>The comparison between <italic>in situ</italic> meteorological data and satellite-derived estimates from the Copernicus platform reveals measurable inconsistencies across the study area. Air temperature differences were generally within &#xb1;0.5&#xb0;C, with localized deviations reaching up to 1.0&#xb0;C in the northern sector. These discrepancies fall within the expected range reported by <xref ref-type="bibr" rid="B40">Li et&#xa0;al. (2013)</xref> and are likely attributable to sharp thermal gradients, variable cloud cover, and atmospheric heterogeneity. Notably, the southern edge of the Intertropical Convergence Zone (ITCZ) was present in the northern portion of the study area, potentially influencing local temperature dynamics.</p>
<p>Wind speed comparisons between <italic>in situ</italic> and satellite-derived data showed more pronounced variability, with differences ranging from &#x2212;2 to +4 m/s. These spatially heterogeneous discrepancies can be attributed to limitations in satellite retrieval techniques&#x2014;particularly scatterometers and altimeters&#x2014;which are sensitive to rain, surface roughness, and land contamination near coastal zones. Additional contributing factors include algorithmic uncertainties, spatial mismatches, and short-term local variability that satellites may fail to capture (<xref ref-type="bibr" rid="B35">Kent et&#xa0;al., 2012</xref>; <xref ref-type="bibr" rid="B51">Mitsopoulos and Pe&#xf1;a, 2023</xref>). Despite these differences in magnitude, wind direction remained consistent across datasets, averaging approximately 225&#xb0;.</p>
<p>Regarding sea surface temperature and sea surface salinity, observed differences ranged from &#x2212;0.6 &#xb0;C to +0.3 &#xb0;C for SST and approximately &#xb1;0.5 units for SSS. Notable anomalies included SST minima of &#x2212;1.0 &#xb0;C in the southern region and SSS maxima of +1.0 in the north. SST discrepancies are primarily due to the depth of <italic>in situ</italic> sensors (&gt;1 m), which measure bulk temperature, whereas satellite sensors detect skin temperature at the surface. Additional sources of error include cloud and atmospheric contamination (e.g., humidity, aerosols, suspended particles), temporal mismatches, diel variation, and differences in spatial resolution (<xref ref-type="bibr" rid="B44">Margaritis et&#xa0;al., 2025</xref>).</p>
<p>Despite the overall coherence between <italic>in situ</italic> and satellite-derived datasets&#x2014;both internally across variables and externally with established literature&#x2014;the presence of localized discrepancies highlights inherent limitations in spatial resolution, sensor depth sensitivity, and retrieval algorithms. These constraints, particularly evident in coastal and dynamically variable regions, underscore the need for cautious interpretation and the continued integration of multi-source observations to enhance the reliability of ocean-atmosphere assessments.</p>
<sec id="s4_1">
<label>4.1</label>
<title>Evidence for the occurrence of El Ni&#xf1;o in 2023-2024</title>
<p>The longest La Ni&#xf1;a event since 1920&#x2014;or at least since 1954&#x2014;was La Ni&#xf1;a 2020&#x2013;2023 (Ormaza-Gonz&#xe1;lez, 2023). The year 2022 ended with a well-established La Ni&#xf1;a phase, and during the first days of January 2023, sea surface temperature (SST) anomalies (&#xb0;C) were recorded as follows: -0.7 in El Ni&#xf1;o 4, -0.8 in 3.4, -0.7 in 3, and -0.8 in 1 + 2 (see <xref ref-type="table" rid="T2"><bold>Table&#xa0;2</bold></xref>). Additionally, the Oceanic Ni&#xf1;o Index (ONI) registered -0.7 &#xb0;C, while the Multivariate El Ni&#xf1;o Index (<xref ref-type="bibr" rid="B47">MEI, 2024</xref>) showed values of -1.1, -0.9, and -0.8 from January through March. Moreover, the Southern Oscillation Index (SOI) exceeded +20 (see <xref ref-type="bibr" rid="B3">BoM, 2025</xref>). These oceanographic and meteorological indicators were strongly coupled and fully consistent with La Ni&#xf1;a conditions, which were expected to persist at least through the first quarter of 2023. During this same period, the Pacific Decadal Oscillation (<xref ref-type="bibr" rid="B60">PDO, 2024</xref>)&#x2014;often described as a long-lived El Ni&#xf1;o-like pattern of Pacific climate variability (<xref ref-type="bibr" rid="B79">Zhang and Levitus, 1997</xref>; <xref ref-type="bibr" rid="B25">Folland et al., 2002</xref>)&#x2014;continued to oscillate around negative values. From January 2019 (-0.07) to July 2025 (-4.00), the PDO reached its lowest value since records began in 1854.</p>
<table-wrap id="T2" position="float">
<label>Table&#xa0;2</label>
<caption>
<p>SST anomalies (C) examples in 2023 for different El Ni&#xf1;o regions. Ni&#xf1;o 1 (far eastern equatorial Pacific): 5&#x2013;10&#xb0;S, 90&#x2013;80&#xb0;W, Ni&#xf1;o 2 (far eastern equatorial Pacific): 0&#x2013;5&#xb0;S, 90&#x2013;80&#xb0;W, Ni&#xf1;o 3 (eastern equatorial Pacific): 5&#xb0;N&#x2013;5&#xb0;S, 150&#x2013;90&#xb0;W, Ni&#xf1;o 3.4 (central equatorial Pacific): 5&#xb0;N&#x2013;5&#xb0;S, 170&#x2013;120&#xb0;W, Ni&#xf1;o 4 (western equatorial Pacific): 5&#xb0;N&#x2013;5&#xb0;S, 160&#xb0;E&#x2013;150&#xb0;W.</p>
</caption>
<table frame="hsides">
<thead>
<tr>
<th valign="middle" align="center">El Ni&#xf1;o region</th>
<th valign="middle" align="center">8 January</th>
<th valign="middle" align="center">6 March</th>
<th valign="middle" align="center">5 June</th>
<th valign="middle" align="center">26 December</th>
</tr>
</thead>
<tbody>
<tr>
<td valign="middle" align="center">4</td>
<td valign="middle" align="center">-0.7</td>
<td valign="middle" align="center">-0.4</td>
<td valign="middle" align="center">0.6</td>
<td valign="middle" align="center">1.4</td>
</tr>
<tr>
<td valign="middle" align="center">3.4</td>
<td valign="middle" align="center">-0.8</td>
<td valign="middle" align="center">-0.2</td>
<td valign="middle" align="center">0.8</td>
<td valign="middle" align="center">2.0</td>
</tr>
<tr>
<td valign="middle" align="center">3</td>
<td valign="middle" align="center">-0.7</td>
<td valign="middle" align="center">0.1</td>
<td valign="middle" align="center">1.1</td>
<td valign="middle" align="center">2.1</td>
</tr>
<tr>
<td valign="middle" align="center">1 + 2</td>
<td valign="middle" align="center">-0.8</td>
<td valign="middle" align="center">1.1</td>
<td valign="middle" align="center">2.3</td>
<td valign="middle" align="center">1.6</td>
</tr>
</tbody>
</table>
</table-wrap>
<p>Beginning in March 2023, rapid surface warming developed along the equatorial Pacific. By the beginning of June sea surface temperature (SST) anomalies in region 3.4 surpass the boundaries of 0.5C, registering 0.8C, from there to the end of December, 2.0&#xb0;C, while in region 1 + 2, 1.6&#xb0;C (<xref ref-type="table" rid="T2"><bold>Table&#xa0;2</bold></xref>). The Oceanic Ni&#xf1;o Index (ONI) registered -2.0 &#xb0;C, and the Southern Oscillation Index (SOI) dropped below -7. These values clearly indicated the onset of a strong El Ni&#xf1;o event, officially declared in September by various agencies including NOAA, the UK Met Office, and the Bureau of Meteorology (BoM). This event had been anticipated to some extent by <xref ref-type="bibr" rid="B58">Ormaza-Gonz&#xe1;lez et&#xa0;al. (2022)</xref>.</p>
<sec id="s4_1_1">
<label>4.1.1</label>
<title>January: 2024 <italic>vs</italic> 1998</title>
<p>The 1997&#x2013;1998 El Ni&#xf1;o event remains the most significant and intense episode recorded since 1950 (<xref ref-type="bibr" rid="B21">Enfield, 2001</xref>), and is widely considered the strongest of the 20th century (<xref ref-type="bibr" rid="B11">Changnon, 2000</xref>; <xref ref-type="bibr" rid="B15">CPC, 2025</xref>). During its peak from October 1997 to January 1998, the Oceanic Ni&#xf1;o Index (ONI) and Multivariate ENSO Index (MEI) reached values between 2.0 and 2.4, averaging 2.2&#xb0;C (<xref ref-type="bibr" rid="B68">Thielen et&#xa0;al., 2023</xref>). Concurrently, in region 1 + 2 and along the western coast of South America, anomalous sea surface temperatures (ASST) exceeded 5&#xb0;C, accompanied by sea level anomalies of approximately 30 cm, with maxima observed in December and January (<xref ref-type="bibr" rid="B8">Carr et&#xa0;al., 2002</xref>). These thermal anomalies, along with SSTs reaching 27&#x2013;28&#xb0;C, suggest the onset of the rainy season, weakening of trade winds, and deepening of the mixed layer and 20&#xb0;C isotherm to depths of at least 50 m and 70 m, respectively. Also, sea level anomalies 20&#x2013;25 cm above normal, and the Intertropical Convergence Zone (ITCZ) shifted southward, positioned at or below the equatorial line.</p>
<p>Conversely, January 2024 exhibited ONI and MEI values averaging 1.9&#xb0;C and 0.90&#xb0;C, respectively, with SSTA in region 1 + 2 ranging from 0.8&#xb0;C to 1.0&#xb0;C (<xref ref-type="bibr" rid="B15">CPC, 2025</xref>). Notably, five downward-propagating Kelvin waves had already reached the eastern Pacific coasts, while an upward Kelvin wave was observed departing from the western Pacific in December.</p>
<p>Meteorological conditions in January 2024 contrasted sharply with those observed during the peak of the 1997&#x2013;1998 El Ni&#xf1;o event. During the latter, winds predominantly blew from the northeast at an average speed of 3.58 &#xb1; 0.83 m/s (<xref ref-type="fig" rid="f11"><bold>Figure&#xa0;11</bold></xref>), reflecting a near-complete collapse of the trade wind system. The Intertropical Convergence Zone (ITCZ) had shifted southward, positioned between 2&#xb0;S and 5&#xb0;S, while SAT across the region consistently exceeded 27 &#xb0;C.</p>
<p>Oppositely, January 2024 recorded an average wind speed of 5.6 &#xb1; 2.0 m/s, with extremes ranging from 0.4 m/s to 11.4 m/s and a predominant wind direction from the southwest (&#x223c;220&#xb0;; <xref ref-type="fig" rid="f11"><bold>Figure&#xa0;11</bold></xref>). Another mark during the 2023&#x2013;2024 event was the intensification of the Pacific anticyclone since September 2023, which led to a reinforcement of the trade winds across the equatorial Pacific. The Southern Pacific Anticyclone, has remained anomalously strong since 2020 (<xref ref-type="bibr" rid="B39">Li et&#xa0;al., 2023</xref>; <xref ref-type="bibr" rid="B80">Zhang et&#xa0;al., 2024</xref>). During this month, the anticyclone exhibited elevated pressure values (&gt;1025 hPa, <xref ref-type="fig" rid="f12"><bold>Figure&#xa0;12</bold></xref>) and an expanded core area relative to climatological norms (<xref ref-type="bibr" rid="B56">Ormaza-Gonz&#xe1;lez, 2024</xref>). This subtropical system plays a critical role in modulating regional precipitation and cloud cover, often suppressing convective activity near its center. Its sustained strength reinforced the trade winds, potentially reflecting a lagged response to prolonged wind forcing or the unprecedented cross-equatorial southerly wind anomalies observed during the 2020&#x2013;2023 triple-dip La Ni&#xf1;a. These anomalies originated in the southeastern tropical Pacific and persisted through January 2024.</p>
<fig id="f11" position="float">
<label>Figure&#xa0;11</label>
<caption>
<p>Wind speed and direction, and SAT during January 1998. Copernicus data, authors reanalysis.</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g011.tif">
<alt-text content-type="machine-generated">Two side-by-side maps show wind speed and direction, and SAT during January 1998. The right map displays sea surface temperatures, with a color gradient from blue to red representing temperatures from 24 to 30 degrees Celsius. Both maps have latitude and longitude coordinates, a north arrow, and a 20-kilometer scale bar.</alt-text>
</graphic></fig>
<fig id="f12" position="float">
<label>Figure&#xa0;12</label>
<caption>
<p>The AP isobaths for January 2024 and average climatology for the month of January. Graphics are constructed with Copernicus Data.</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g012.tif">
<alt-text content-type="machine-generated">Atmospheric Isobars, showing the South Pacific Anticyclone. The left map shows January 2024 pressures, with contours colored from dark blue to red, ranging from eight hundred to one thousand hectopascals. The right map depicts climatological data with similar gradient colors and pressure values, highlighting differences between the two periods. Contour lines illustrate pressure variations across the continent.</alt-text>
</graphic></fig>
<p>Meanwhile, the Intertropical Convergence Zone (ITCZ) experienced a notable northward displacement, with its core firmly positioned between 2&#xb0;N and 3&#xb0;N &#x2014;significantly farther north than its climatological average for this time of year (<xref ref-type="bibr" rid="B57">Ormaza-Gonz&#xe1;lez and Cede&#xf1;o, 2017</xref>). This shift was accompanied by a strengthened trade wind system, although its spatial and directional characteristics differed from those observed during the 1998 El Ni&#xf1;o event. Seasonal rainfall intensity during El Ni&#xf1;o or La Ni&#xf1;a episodes is influenced not only by sea surface temperatures (SSTs) but also by the latitudinal position of the ITCZ (<xref ref-type="bibr" rid="B24">Fahrin et&#xa0;al., 2024</xref>). In January 2024, despite SSTs exceeding 27 &#xb0;C along the Ecuadorian coast (approximately 1.8&#xb0;N to 2.5&#xb0;S), rainfall was markedly below average. For instance, Guayaquil&#x2014;a key reference city for assessing El Ni&#xf1;o impacts&#x2014;recorded only 114 mm of rainfall, well below its typical January average of over 200 mm. This rainfall deficit is attributed to the ITCZ&#x2019;s anomalously northern position, which limited convective activity over the region. In contrast, during the strong El Ni&#xf1;o events of 1982&#x2013;1983 and 1997&#x2013;1998, Guayaquil received 6210 mm and 5990 mm of rainfall, respectively (see, <xref ref-type="bibr" rid="B22">ERAS 2025</xref>), whereas the 2023&#x2013;2024 event yielded only 1340 mm for the entire season. These observations underscore the distinct atmospheric dynamics of the 2023&#x2013;2024 event, highlighting its divergence from canonical El Ni&#xf1;o patterns.</p>
<p>The surface air temperature in 1998 was also higher than that registered in 2024; 27.18 &#xb1; 0.13 and 26.53 &#xb1; 0.87 in the same order, with maxima of 27.18 and 28.60; and minima of 26.83 and 24.30, respectively. On average they were similar, but ranges were dissimilar in about 0.35 and 4.3 C making air masses consistently higher SAT in 1998, therefore enhancing humidity and precipitation (<xref ref-type="bibr" rid="B45">Marsh, 2011</xref>), which did not occur in 2024.</p>
<p>Sea surface temperature (SST) patterns further underscored the divergence between the 1998 and 2024 conditions. In January 1998, SSTs decreased latitudinally from south to north, ranging from 28.02&#xb0;C to 27.69&#xb0;C, with a regional average of 27.90 &#xb1; 0.11 &#xb0;C (<xref ref-type="fig" rid="f13"><bold>Figure&#xa0;13</bold></xref>). During the development of the 2023&#x2013;2024 El Ni&#xf1;o, <xref ref-type="bibr" rid="B65">Saltos-Aguilar and Ormaza-Gonz&#xe1;lez (2023)</xref>; <xref ref-type="bibr" rid="B14">CNP (2023)</xref> reported observational data from monthly cruises conducted along the entire Ecuadorian coast, spanning approximately 3&#xb0;S to the equator&#x2014;within region 1 + 2 (90&#xb0;W&#x2013;80&#xb0;W, 0&#xb0;&#x2013;10&#xb0;S). SST measurements in the southern sector (Gulf of Guayaquil) and northern sector were as follows:</p>
<fig id="f13" position="float">
<label>Figure&#xa0;13</label>
<caption>
<p>SSS and SST distribution of January monthly averaged in the area researched during January 1998.</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g013.tif">
<alt-text content-type="machine-generated">Two maps depict sea surface temperatures along a coastal region in different color gradients. The left map shows higher temperatures, ranging from 30 to 34 degrees Celsius, indicated by green and turquoise shades. The right map shows lower temperatures, from 25 to 30 degrees Celsius, in yellow to light green shades. Red dots mark specific measured points with temperature labels. A north arrow and scale bars are included.</alt-text>
</graphic></fig>
<list list-type="bullet">
<list-item>
<p>July: 27.8&#xb0;C in both regions</p></list-item>
<list-item>
<p>August: 25.6&#xb0;C (south), 27.2&#xb0;C (north)</p></list-item>
<list-item>
<p>September: 25.8&#xb0;C (south), 25.2&#xb0;C (north)</p></list-item>
</list>
<p>In January 2024, SSTs exhibited a broader latitudinal gradient, ranging from 25.90 &#xb0;C in the south to 28.60 &#xb0;C in the north, with a slightly lower regional average of 27.49 &#xb1; 0.63 &#xb0;C. This reversal in the latitudinal SST gradient, along with increased variability, supports the presence of relatively stronger trade winds and altered ocean-atmosphere coupling during the 2024 event. Moreover, SST distribution and values in 2024 closely resembled climatological patterns (<xref ref-type="fig" rid="f4"><bold>Figure&#xa0;4</bold></xref>), suggesting a return to seasonal norms.</p>
<p>Sea surface salinity (SSS, <xref ref-type="fig" rid="f13"><bold>Figure&#xa0;13</bold></xref>) in January 1998 showed a slight increase from north to south, ranging from 30.73 to 31.34, with an average of 31.07 &#xb1; 0.18. In contrast, SSS in January 2024 was notably higher, increasing from 33 to 34 across the same latitudinal gradient. The horizontal distribution of isohaline contours was conspicuously different. Specifically, low-salinity water masses originating from Panam&#xe1; Bay (<xref ref-type="bibr" rid="B2">Alory et al., 2012</xref>) appeared to intrude southward into regions typically characterized by higher salinity. This pattern mirrored the climatological SSS distribution (<xref ref-type="fig" rid="f4"><bold>Figure&#xa0;4</bold></xref>), further indicating that January 2024 reflected a seasonal configuration rather than a pronounced El Ni&#xf1;o signal.</p>
<p>Taken together, these variables suggest that oceanographic conditions in early 2024 were more consistent with climatological expectations, and no strong impact from the 2023&#x2013;2024 El Ni&#xf1;o event was evident.</p>
</sec>
</sec>
<sec id="s4_2">
<label>4.2</label>
<title>Mixed layer depth and 20 C isotherm variability</title>
<p>The depth of the thermocline, or mixed layer, typically increases during El Ni&#xf1;o events, often beginning in December or even November. Historical records from the 1982&#x2013;1983 and 1997&#x2013;1998 El Ni&#xf1;o episodes indicate mixed layer depths (MLD) exceeding 100 m (<xref ref-type="bibr" rid="B43">Mangum et&#xa0;al., 1986</xref>; <xref ref-type="bibr" rid="B17">Cucalon, 1987</xref>; <xref ref-type="bibr" rid="B21">Enfield, 2001</xref>), suggesting strong surface stratification during periods of pronounced impact along the Ecuadorian and Peruvian coasts. Notably, in those events, the deepening of the MLD and the 20 &#xb0;C isotherm began later in the preceding year.</p>
<p>In contrast, during the 2023&#x2013;2024 El Ni&#xf1;o, <italic>in situ</italic> measurements taken through November offshore the Ecuadorian coast revealed MLD values ranging from 10 to 14 m between 2.5&#xb0;S and 0.5&#xb0;N along 81&#xb0;W longitude, corresponding to area 1 + 2. By January 2024, the spatial distribution of the MLD showed partial alignment with the January climatology (1993&#x2013;2022), though it was generally deeper than the climatological range of 13 to 19 m (<xref ref-type="fig" rid="f14"><bold>Figure&#xa0;14</bold></xref>). A prominent nucleus centered near 0.5&#xb0;S, between 81.0&#xb0; and 80.5&#xb0;W, was observed in 2024 but is absent from the climatological pattern. The northern core around 1&#xb0;N appears in both datasets, though it was deeper in 2024. Overall, the January 2024 thermocline was deeper than the climatological average, except in the southern region where it was comparatively shallower. By February 2024, the MLD had shoaled by approximately 7 m near 2&#xb0;S (<xref ref-type="bibr" rid="B6">Bustos O&#xf1;a et&#xa0;al., 2024</xref>), supporting the conclusion that the southward influence of the Humboldt Current effectively blocked the intrusion of warm waters from Panam&#xe1; Bay.</p>
<fig id="f14" position="float">
<label>Figure&#xa0;14</label>
<caption>
<p>Spatial distribution of the mixed layer and 20C vertical isotherm depth climatology from 1993 to 2022.</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g014.tif">
<alt-text content-type="machine-generated">Two contour maps depict different oceanographic variables over a region. The left map shows temperature distributions, ranging from blue (cooler) to green (warmer), while the right map illustrates depth variations with colors from orange (shallower) to green (deeper). Latitude and longitude are marked, and a scale indicates distance. A color bar on the right shows depth in meters.</alt-text>
</graphic></fig>
<p>The depth and spatial distribution of the 20 &#xb0;C isotherm revealed notable differences among the climatology, 1998, and 2024 datasets, both in absolute values and isoline configuration. The climatological pattern features isolines between 30 and 42 m depth (<xref ref-type="fig" rid="f15"><bold>Figure&#xa0;15</bold></xref>), generally aligned parallel to the coastline. In January 1998 (<xref ref-type="fig" rid="f15"><bold>Figure&#xa0;15</bold></xref>), the isotherm extended offshore to depths of up to 115 m, with shallower values around 50 m near the coast. This configuration suggests a horizontal thermal gradient indicative of reduced vertical mixing, increased stratification with distance from land (<xref ref-type="bibr" rid="B21">Enfield, 2001</xref>), thermocline tilt, regional circulation features, and the influence of broader oceanic processes across the central-eastern equatorial Pacific.</p>
<fig id="f15" position="float">
<label>Figure&#xa0;15</label>
<caption>
<p>Spatial distribution of the mixed layer and 20C vertical isotherm depth during January 1998.</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g015.tif">
<alt-text content-type="machine-generated">Two contour maps side by side. The left map shows varying altitudes in shades of blue to green. The right map represents elevations in shades of green to orange, indicating higher altitudes. Both maps display latitude and longitude axes with a scale bar and a compass rose. Each map includes a color scale indicating meters.</alt-text>
</graphic></fig>
<p>In contrast, January 2024 exhibited two distinct nuclei of maximum depth&#x2014;approximately 55 m&#x2014;located in the southern and northern portions of the study area. These features formed a wave-like pattern, consistent with the spatial distribution observed in the MLD. Despite the presence of a subsurface thermal nucleus&#x2014;which may indicate localized upwelling as surface waters moved northward near 1&#xb0;S&#x2014;the overall vertical structure of the water column during the cruise reflected seasonal conditions rather than anomalous El Ni&#xf1;o-driven dynamics.</p>
<p>Furthermore, the subsurface salinity and temperature profiles support the hypothesis that, at the time of observation, the strong 2023&#x2013;2024 El Ni&#xf1;o did not exert a measurable impact on coastal subsurface stratification. Consequently, both sea surface temperature (SST) and sea surface salinity (SSS) suggest the absence of a significant El Ni&#xf1;o signal along the Ecuadorian coast during early 2024.</p>
</sec>
<sec id="s4_3">
<label>4.3</label>
<title>Vertical distribution and water masses</title>
<p>Oceanographic and meteorological variability in the study region during January 2024 strongly suggest that surface conditions were predominantly influenced by the seasonal intrusion of Panam&#xe1; Bay waters from the north and the counteracting presence of the Humboldt Current from the south. These opposing forces establish a dynamic convergence zone between the equator and northern Ecuador.</p>
<p>Hydrographic data indicate that Panam&#xe1; Bay water&#x2014;extending longitudinally from 85&#xb0;W to 90&#xb0;W and latitudinally from 5&#xb0;N to 5&#xb0;S&#x2014;is characterized by sea surface salinities (SSS) below 33 and sea surface temperatures (SST) exceeding 28 &#xb0;C. This water mass was observed in the northern portion of the study area, penetrating southward to approximately the equatorial line.</p>
<p>Deeper water masses, such as Antarctic Intermediate Water (AAIW), were detected as far north as approximately 2&#xb0;N. According to <xref ref-type="bibr" rid="B4">Bostock et&#xa0;al. (2013)</xref>, this corresponds to the South Pacific subtype, identified at around 1000 m depth. Temperature-salinity (T-S) correlations indicate a density exceeding 1.027 g/cm&#xb3;. The primary drivers of AAIW presence in these latitudes include:</p>
<p>1. Meridional Overturning Circulation (MOC). The AAIW is part of the global thermohaline circulation, which moves water masses across vast distances. After forming near the Antarctic Polar Front, AAIW travels northward along western boundary currents and interior pathways, eventually crossing the equator.</p>
<p>2. Western Boundary Currents.</p>
<p>In the South Pacific, AAIW is advected northward via the Peru-Chile Undercurrent and the Equatorial Undercurrent system. These currents help funnel intermediate waters toward the equator and beyond.</p>
<p>3. Equatorial Upwelling and Mixing.</p>
<p>Near the equator, strong upwelling and vertical mixing can bring intermediate waters closer to the surface. This mixing allows AAIW to influence nutrient and carbon dynamics in equatorial regions.</p>
<p>4. Diapycnal and Isopycnal Mixing.</p>
<p>Turbulent mixing across density surfaces (diapycnal) and along them (isopycnal) helps redistribute AAIW across latitudes. These processes are especially active in the equatorial Pacific due to high energy from wind and tidal forces.</p>
<p>5. Topographic Steering.</p>
<p>The complex bathymetry of the eastern Pacific, including ridges and continental slopes, can steer AAIW northward. These features guide water masses along preferred pathways, sometimes funneling them into equatorial zones.</p>
<p>Deeper, colder, and nutrient-rich water masses&#x2014;and the dynamics that govern them&#x2014;are essential to understanding ENSO-driven variability in deep ocean circulation across the Southeast Pacific. Recent findings by <xref ref-type="bibr" rid="B69">Torres-Godoy et&#xa0;al. (2025)</xref> demonstrate that interannual fluctuations in the transport of Antarctic Intermediate Water (AAIW) are intricately linked to ENSO phases, highlighting a dynamic coupling between intermediate-depth processes and surface climate variability. Complementary research by <xref ref-type="bibr" rid="B74">Wang et&#xa0;al. (2023)</xref> examined the combined influence of the Southern Annular Mode (SAM) and ENSO on Antarctic sea ice, suggesting that variations in AAIW formation&#x2014;modulated by these climate modes&#x2014;can extend their impact into tropical latitudes, thereby shaping surface ocean conditions. Furthermore, <xref ref-type="bibr" rid="B19">Dou and Zhang (2023)</xref> reported a weakening in the relationship between ENSO and Antarctic sea ice, and by extension AAIW, over recent decades, indicating a potential shift in the mechanisms that connect deep ocean circulation with surface climate dynamics.</p>
</sec>
<sec id="s4_4">
<label>4.4</label>
<title>Chlorophyll-a and surface currents</title>
<p>Further evidence that the 2023&#x2013;2024 El Ni&#xf1;o event did not produce significant coastal impacts in the study region comes from the analysis of Chlorophyll-a (Cl-a) concentrations, a key indicator of primary productivity and the foundation of the marine trophic chain. Cl-a data was used alongside surface current observations to assess oceanographic conditions. In tropical regions, Cl-a variability is closely tied to physical processes: elevated concentrations typically result from cold, nutrient-rich upwelling, while reduced levels are associated with warm, oligotrophic waters depleted in essential nutrients such as nitrogen, phosphorus, silicate, and trace micronutrients.</p>
<p>The productivity of Ecuadorian coastal waters is primarily governed by the seasonal dominance of northward and southward currents, which prevail from June to November and December to May, respectively (<xref ref-type="bibr" rid="B13">Chinacalle-Mart&#xed;nez et&#xa0;al., 2021</xref>). While El Ni&#xf1;o and La Ni&#xf1;a events are known to exert significant influence on trophic dynamics, neutral or seasonally driven conditions tend to maintain a stable trophic structure.</p>
<p>In January 2024, satellite-derived Cl-a concentrations were notably higher than those recorded in January 2023, despite the latter occurring under La Ni&#xf1;a conditions (<xref ref-type="fig" rid="f16"><bold>Figure&#xa0;16</bold></xref>). This finding is consistent with previous hydrographic and atmospheric observations and is further supported by surface current data from Copernicus satellite products, which indicate a predominantly northward flow during January 2024&#x2014;mirroring the circulation patterns observed throughout the 2020&#x2013;2023 triple-dip La Ni&#xf1;a period. .</p>
<fig id="f16" position="float">
<label>Figure&#xa0;16</label>
<caption>
<p>Chlorophyll-a content during January 2023 (left) and 2024 (right).</p>
</caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fmars-12-1737039-g016.tif">
<alt-text content-type="machine-generated">Two maps display chlorophyll concentration off the coast of a region. The left map shows predominantly green areas indicating lower concentration levels, while the right map has yellow to red areas near the coast representing higher concentration levels. A color scale on the right quantifies the concentration from 0.00 to 2.27 mg per cubic meter.</alt-text>
</graphic></fig>
</sec>
</sec>
<sec id="s5" sec-type="conclusions">
<label>5</label>
<title>Conclusions</title>
<p>The comparative analysis between <italic>in situ</italic> meteorological and oceanographic measurements and Copernicus satellite estimates demonstrates that both sources are broadly reliable, though each carries inherent limitations. Localized discrepancies in air temperature, wind speed, sea surface temperature, and salinity highlight the indispensable role of sustained, high&#x2212;frequency <italic>in situ</italic> sampling, particularly in regions shaped by sharp and dynamic gradients and the Intertropical Convergence Zone.</p>
<p>These findings underscore the complementary nature of field and remote sensing approaches. <italic>In situ</italic> datasets provide essential validation and context for satellite observations, while satellites extend spatial coverage and enable long&#x2212;term monitoring. The integration of these methods enhances confidence in ocean&#x2212;atmosphere assessments and strengthens predictive frameworks for climate variability in the eastern equatorial Pacific.</p>
<p>Despite its classification as a strong El Ni&#xf1;o, the 2023&#x2013;2024 event did not produce the expected coastal impacts along the eastern Pacific. Subsurface thermal and salinity structures remained within seasonal norms, and the mixed layer depth and 20 &#xb0;C isotherm showed no signs of the anomalous deepening typically associated with canonical El Ni&#xf1;o events. The intensification of the Pacific anticyclone and reinforced trade winds since September 2023 further suggest atypical atmospheric dynamics.</p>
<p>Oceanographic observations confirm that local processes dominated. Mixed layers remained shallow, salinity and temperature gradients reflected Panam&#xe1; Bay intrusions and Humboldt Current influence, and no basin&#x2212;wide anomalies were detected. A particularly novel finding was the serendipitous detection of Antarctic Intermediate Water (AAIW) as far north as 2&#xb0;N. This highlights the importance of deep vertical profiling of temperature and salinity, an area often overlooked in ENSO studies that focus primarily on surface variability. The observed T/S relationships underscore how intermediate waters connect high&#x2212;latitude climate models with tropical ocean conditions, revealing the complexity of deep ocean&#x2013;surface climate linkages in the Southeast Pacific.</p>
<p>Biological and meteorological indicators reinforce this divergence. Chlorophyll&#x2212;a concentrations and surface current patterns in January 2024 resembled those typical of neutral or La Ni&#xf1;a conditions, likely reflecting residual influence from the 2020&#x2013;2023 triple&#x2212;dip La Ni&#xf1;a. Rainfall also remained near seasonal averages: in Guayaquil, January&#x2013;April totals were ~1340&#x2013;1500 mm, far below the ~6000 mm recorded during the extreme 1982&#x2013;1983 and 1997&#x2013;1998 El Ni&#xf1;o events. Although the northern coast experienced above&#x2212;average precipitation, it did not approach the extreme levels characteristic of strong eastern Pacific El Ni&#xf1;o events.</p>
<p>A rigorous, multi&#x2212;layered approach&#x2014;integrating meteorology, oceanographic <italic>in situ</italic> surface observations, satellite monitoring, and deep vertical profiling&#x2014;provides the most reliable pathway to capture this complexity. Establishing such epistemic clarity is essential for anticipating impacts, informing policy, and safeguarding coastal societies that depend on accurate knowledge of ocean dynamics.</p>
</sec>
</body>
<back>
<sec id="s6" sec-type="data-availability">
<title>Data availability statement</title>
<p>The original contributions presented in the study are included in the article/<xref ref-type="supplementary-material" rid="SM1"><bold>Supplementary Material</bold></xref>. Further inquiries can be directed to the corresponding author.</p></sec>
<sec id="s7" sec-type="author-contributions">
<title>Author contributions</title>
<p>FO-G: Conceptualization, Methodology, Formal Analysis, Writing &#x2013; review &amp; editing, Supervision, Validation, Writing &#x2013; original draft, Investigation. CM-B: Visualization, Data curation, Software, Resources, Writing &#x2013; review &amp; editing. EE: Writing &#x2013; review &amp; editing, Software, Data curation, Visualization, Formal Analysis. J-NP: Writing &#x2013; review &amp; editing, Resources, Funding acquisition, Project administration. FM: Supervision, Writing &#x2013; review &amp; editing, Resources, Funding acquisition, Project administration. J-FL: Data curation, Formal Analysis, Software, Resources, Investigation, Writing &#x2013; review &amp; editing.</p></sec>
<ack>
<title>Acknowledgments</title>
<p>We gratefully acknowledge the reviewers for their supportive, encouraging, and constructive comments, which greatly improved the quality of this manuscript. Data were acquired in 2024 during the IRD-CNRS oceanographic campaign SUPER-MOUV on board the French oceanographic research vessel Pourquoi Pas?. This work was carried out in the framework of the collaboration between the CNRS-INSU, IRD, IFREMER and ESPOL, Nice, Rennes, Antilles and Sorbonne Universities, and ANR Fluid2Slip (Audrey Galv&#xe9;). We thank the captains (Philippe Moineaux and Pierre-Yves Le Maer) and the crews of R/V Pourquoi pas? for their efficient work. Thanks to the French Embassies in Ecuador and in Colombia and INOCAR for their help before and during the cruise. We thank the SUPER-MOUV Science Party for their help in gathering and processing the data during the campaign and for the fruitful scientific discussions onboard and at different stages afterwards. We also thank the Maritime Faculty of ESPOL (FIMCM-ESPOL), for the support in different ways.</p>
</ack>
<sec id="s9" sec-type="COI-statement">
<title>Conflict of interest</title>
<p>The authors declared that this work was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest.</p></sec>
<sec id="s10" sec-type="ai-statement">
<title>Generative AI statement</title>
<p>The author(s) declare that Generative AI was not used in the creation of this manuscript.</p>
<p>Any alternative text (alt text) provided alongside figures in this article has been generated by Frontiers with the support of artificial intelligence and reasonable efforts have been made to ensure accuracy, including review by the authors wherever possible. If you identify any issues, please contact us.</p></sec>
<sec id="s11" sec-type="disclaimer">
<title>Publisher&#x2019;s note</title>
<p>All claims expressed in this article are solely those of the authors and do not necessarily represent those of their affiliated organizations, or those of the publisher, the editors and the reviewers. Any product that may be evaluated in this article, or claim that may be made by its manufacturer, is not guaranteed or endorsed by the publisher.</p></sec>
<sec id="s12" sec-type="supplementary-material">
<title>Supplementary material</title>
<p>The Supplementary Material for this article can be found online at: <ext-link ext-link-type="uri" xlink:href="https://www.frontiersin.org/articles/10.3389/fmars.2025.1737039/full#supplementary-material">https://www.frontiersin.org/articles/10.3389/fmars.2025.1737039/full#supplementary-material</ext-link></p>
<supplementary-material xlink:href="DataSheet1.pdf" id="SM1" mimetype="application/pdf"/>
<supplementary-material xlink:href="DataSheet2.pdf" id="SM2" mimetype="application/pdf"/></sec>
<ref-list>
<title>References</title>
<ref id="B1">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Adamson</surname> <given-names>G.</given-names></name>
</person-group> (<year>2022</year>). 
<article-title>El Ni&#xf1;o without &#x2018;El Ni&#xf1;o&#x2019;? Path dependency and the definition problem in El Ni&#xf1;o Southern Oscillation research</article-title>. <source>Environ. Plann. E.: Nat. Sp.</source> <volume>6</volume>, <fpage>2047</fpage>&#x2013;<lpage>2070</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1177/25148486221120546</pub-id>
</mixed-citation>
</ref>
<ref id="B2">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Alory</surname> <given-names>G.</given-names></name>
<name><surname>Maes</surname> <given-names>C.</given-names></name>
<name><surname>Delcroix</surname> <given-names>T.</given-names></name>
<name><surname>Reul</surname> <given-names>N.</given-names></name>
<name><surname>Illig</surname> <given-names>S.</given-names></name>
</person-group> (<year>2012</year>). 
<article-title>Seasonal dynamics of sea surface salinity off Panama: The far Eastern Pacific Fresh Pool</article-title>. <source>J. Geophys. Res.: Ocean.</source> <volume>117</volume>, <elocation-id>C04028</elocation-id>. doi:&#xa0;<pub-id pub-id-type="doi">10.1029/2011JC007802</pub-id>
</mixed-citation>
</ref>
<ref id="B3">
<mixed-citation publication-type="web">
<person-group person-group-type="author"><collab>BoM</collab>
</person-group> (<year>2025</year>). Available online at: <uri xlink:href="http://www.bom.gov.au/climate/enso">http://www.bom.gov.au/climate/enso</uri> (Accessed <date-in-citation content-type="access-date">September 30, 2025</date-in-citation>).
</mixed-citation>
</ref>
<ref id="B4">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Bostock</surname> <given-names>H. C.</given-names></name>
<name><surname>Sutton</surname> <given-names>P. J.</given-names></name>
<name><surname>Williams</surname> <given-names>M. J.</given-names></name>
<name><surname>Opdyke</surname> <given-names>B. N.</given-names></name>
</person-group> (<year>2013</year>). 
<article-title>Reviewing the circulation and mixing of Antarctic Intermediate Water in the South Pacific using evidence from geochemical tracers and Argo float trajectories</article-title>. <source>Dp. Sea. Res. Part I.: Oceanogr. Res. Pap.</source> <volume>73</volume>, <fpage>84</fpage>&#x2013;<lpage>98</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1016/j.dsr.2012.11.007</pub-id>
</mixed-citation>
</ref>
<ref id="B5">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Brugnoli</surname> <given-names>E.</given-names></name>
<name><surname>Molina</surname> <given-names>L.</given-names></name>
<name><surname>Till</surname> <given-names>I.</given-names></name>
<name><surname>Morales-Ram&#xed;rez</surname> <given-names>A.</given-names></name>
<name><surname>D&#xed;az-Ferguson</surname> <given-names>E.</given-names></name>
</person-group> (<year>2023</year>). 
<article-title>Oceanographic conditions and mesozooplankton in the North zone of Coiba National Park (Eastern Tropical Pacific), Panama, Central America</article-title>. <source>Reg. Stud. Mar. Sci.</source> <volume>66</volume>, <elocation-id>103136</elocation-id>. doi:&#xa0;<pub-id pub-id-type="doi">10.1016/j.rsma.2023.103136</pub-id>
</mixed-citation>
</ref>
<ref id="B6">
<mixed-citation publication-type="web">
<person-group person-group-type="author">
<name><surname>Bustos O&#xf1;a</surname> <given-names>A.</given-names></name>
<name><surname>Ponce Villao</surname> <given-names>G.</given-names></name>
<name><surname>C&#xe1;rdenas Condoy</surname> <given-names>J.</given-names></name>
<name><surname>Ormaza-Gonz&#xe1;les</surname> <given-names>F.</given-names></name>
</person-group> (<year>2024</year>). <source>Condiciones Oceanogr&#xe1;ficas costeras durante febrero 2024</source>. Available online at: <uri xlink:href="https://camaradepesqueria.ec/condiciones-oceanograficas-costeras-durante-febrero-2024/">https://camaradepesqueria.ec/condiciones-oceanograficas-costeras-durante-febrero-2024/</uri> (Accessed <date-in-citation content-type="access-date">September 12, 2025</date-in-citation>).
</mixed-citation>
</ref>
<ref id="B7">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Cai</surname> <given-names>W.</given-names></name>
<name><surname>Santoso</surname> <given-names>A.</given-names></name>
<name><surname>Wang</surname> <given-names>G.</given-names></name>
<name><surname>Yeh</surname> <given-names>S. W.</given-names></name>
<name><surname>An</surname> <given-names>S. I.</given-names></name>
<name><surname>Cobb</surname> <given-names>K. M.</given-names></name>
<etal/>
</person-group>. (<year>2015</year>). 
<article-title>ENSO and greenhouse warming</article-title>. <source>Nat. Climate Change</source> <volume>5</volume>, <fpage>849</fpage>&#x2013;<lpage>859</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1038/NCLIMATE2743</pub-id>
</mixed-citation>
</ref>
<ref id="B8">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Carr</surname> <given-names>M. E.</given-names></name>
<name><surname>Strub</surname> <given-names>P. T.</given-names></name>
<name><surname>Thomas</surname> <given-names>A. C.</given-names></name>
<name><surname>Blanco</surname> <given-names>J. L.</given-names></name>
</person-group> (<year>2002</year>). 
<article-title>Evolution of 1996&#x2013;1999 La Ni&#xf1;a and El Ni&#xf1;o conditions off the western coast of South America: a remote sensing perspective</article-title>. <source>J. Geophys. Res.: Ocean.</source> <volume>107</volume>, <fpage>29</fpage>&#x2013;<lpage>21</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1029/2001JC001183</pub-id>
</mixed-citation>
</ref>
<ref id="B9">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Chaigneau</surname> <given-names>A.</given-names></name>
<name><surname>Abarca Del Rio</surname> <given-names>R.</given-names></name>
<name><surname>Colas</surname> <given-names>F.</given-names></name>
</person-group> (<year>2006</year>). 
<article-title>Lagrangian study of the Panama Bight and surrounding regions</article-title>. <source>J. Geophys. Res.: Ocean.</source> <volume>111</volume>, <fpage>C09013</fpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1029/2006JC003530</pub-id>
</mixed-citation>
</ref>
<ref id="B10">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Chaigneau</surname> <given-names>A.</given-names></name>
<name><surname>Dominguez</surname> <given-names>N.</given-names></name>
<name><surname>Eldin</surname> <given-names>G.</given-names></name>
<name><surname>Vasquez</surname> <given-names>L.</given-names></name>
<name><surname>Flores</surname> <given-names>R.</given-names></name>
<name><surname>Grados</surname> <given-names>C.</given-names></name>
<etal/>
</person-group>. (<year>2013</year>). 
<article-title>Near-coastal circulation in the Northern Humboldt Current System from shipboard ADCP data</article-title>. <source>J. Geophys. Res.: Ocean.</source> <volume>118</volume>, <fpage>5251</fpage>&#x2013;<lpage>5266</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1002/jgrc.20328</pub-id>
</mixed-citation>
</ref>
<ref id="B11">
<mixed-citation publication-type="book">
<person-group person-group-type="editor">
<name><surname>Changnon</surname> <given-names>S. A.</given-names></name>
</person-group> (Ed.) (<year>2000</year>). <source>El Ni&#xf1;o 1997-1998: the climate event of the century</source> (<publisher-loc>Oxford, England</publisher-loc>: 
<publisher-name>Oxford University Press</publisher-name>).
</mixed-citation>
</ref>
<ref id="B12">
<mixed-citation publication-type="book">
<person-group person-group-type="author">
<name><surname>Charcot</surname> <given-names>J.</given-names></name>
</person-group> (<year>1910</year>). &#x201c;
<article-title>Story of the &#x2018;Pourquoi-Pas&#x2019;</article-title>,&#x201d; in <source>SA Supplements</source>, vol. <volume>69</volume>. , <fpage>278</fpage>. Available online at: <uri xlink:href="https://www.scientificamerican.com/issue/supplements/1910/04-30/">https://www.scientificamerican.com/issue/supplements/1910/04-30/</uri> (Accessed <date-in-citation content-type="access-date">December 15, 2025</date-in-citation>).
</mixed-citation>
</ref>
<ref id="B13">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Chinacalle-Mart&#xed;nez</surname> <given-names>N.</given-names></name>
<name><surname>Garc&#xed;a-Rada</surname> <given-names>E.</given-names></name>
<name><surname>L&#xf3;pez-Mac&#xed;as</surname> <given-names>J.</given-names></name>
<name><surname>Pinoargote</surname> <given-names>S.</given-names></name>
<name><surname>Loor</surname> <given-names>G.</given-names></name>
<name><surname>Zevallos-Rosado</surname> <given-names>J.</given-names></name>
<etal/>
</person-group>. (<year>2021</year>). 
<article-title>Oceanic primary production trend patterns along coast of Ecuador</article-title>. <source>Neotrop. Biodiv.</source> <volume>7</volume>, <fpage>379</fpage>&#x2013;<lpage>391</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1080/23766808.2021.1964915</pub-id>
</mixed-citation>
</ref>
<ref id="B14">
<mixed-citation publication-type="web">
<person-group person-group-type="author"><collab>CNP</collab>
</person-group> (<year>2023</year>). <source>Coastal-oceanographic information</source>. Available online at: <uri xlink:href="https://camaradepesqueria.ec/condiciones-oceanograficas-durante-septiembre-2023/">https://camaradepesqueria.ec/condiciones-oceanograficas-durante-septiembre-2023/</uri> ().
</mixed-citation>
</ref>
<ref id="B15">
<mixed-citation publication-type="web">
<person-group person-group-type="author"><collab>CPC</collab>
</person-group> (<year>2025</year>). <source>Cold &amp; Warm Episodes by Season</source>. Available online at: <uri xlink:href="https://origin.cpc.ncep.noaa.gov/products/analysis_monitoring/ensostuff/ONI_v5.php">https://origin.cpc.ncep.noaa.gov/products/analysis_monitoring/ensostuff/ONI_v5.php</uri> (Accessed <date-in-citation content-type="access-date">September 4, 2025</date-in-citation>).
</mixed-citation>
</ref>
<ref id="B16">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Crawford</surname> <given-names>G.</given-names></name>
<name><surname>Mepstead</surname> <given-names>M.</given-names></name>
<name><surname>D&#xed;az-Ferguson</surname> <given-names>E.</given-names></name>
</person-group> (<year>2024</year>). 
<article-title>Characterizing oceanographic conditions near Coiba Island and Pacific Panama using 20 years of satellite-based wind stress, SST and chlorophyll-a measurements</article-title>. <source>Mar. Fish. Sci. (MAFIS)</source> <volume>37</volume>, <fpage>391</fpage>&#x2013;<lpage>411</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.47193/mafis.37X2024010112</pub-id>
</mixed-citation>
</ref>
<ref id="B17">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Cucalon</surname> <given-names>E.</given-names></name>
</person-group> (<year>1987</year>). 
<article-title>Oceanographic variability off Ecuador associated with an El Ni&#xf1;o event in 1982&#x2013;1983</article-title>. <source>J. Geophys. Res.</source> <volume>92</volume>, <fpage>14309</fpage>&#x2013;<lpage>14322</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1029/JC092iC13p14309</pub-id>
</mixed-citation>
</ref>
<ref id="B18">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>D&#x2019;Croz.</surname> <given-names>L.</given-names></name>
<name><surname>Del Rosario</surname> <given-names>J. B.</given-names></name>
<name><surname>G&#xf3;mez</surname> <given-names>J. A.</given-names></name>
</person-group> (<year>1991</year>). 
<article-title>Upwelling and phytoplankton in the Bay of Panama</article-title>. <source>Rev. Biol Trop.</source> <volume>39</volume>, <fpage>233</fpage>&#x2013;<lpage>241</lpage>. Available online at: <uri xlink:href="https://archivo.revistas.ucr.ac.cr/index.php/rbt/article/view/24866">https://archivo.revistas.ucr.ac.cr/index.php/rbt/article/view/24866</uri> (Accessed <date-in-citation content-type="access-date">August 7, 2025</date-in-citation>).
</mixed-citation>
</ref>
<ref id="B19">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Dou</surname> <given-names>J.</given-names></name>
<name><surname>Zhang</surname> <given-names>R.</given-names></name>
</person-group> (<year>2023</year>). 
<article-title>Weakened relationship between ENSO and Antarctic sea ice in recent decades</article-title>. <source>Climate Dynamics</source> <volume>60</volume>, <fpage>1313</fpage>&#x2013;<lpage>1327</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1007/s00382-022-06364-4</pub-id>
</mixed-citation>
</ref>
<ref id="B20">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Echevin</surname> <given-names>V.</given-names></name>
<name><surname>Colas</surname> <given-names>F.</given-names></name>
<name><surname>Espinoza-Morriberon</surname> <given-names>D.</given-names></name>
<name><surname>Vasquez</surname> <given-names>L.</given-names></name>
<name><surname>Anculle</surname> <given-names>T.</given-names></name>
<name><surname>Gutierrez</surname> <given-names>D.</given-names></name>
</person-group> (<year>2018</year>). 
<article-title>Forcings and evolution of the 2017 coastal el Nino off northern Peru and Ecuador</article-title>. <source>Front. Mar. Sci.</source> <volume>5</volume>. doi:&#xa0;<pub-id pub-id-type="doi">10.3389/fmars.2018.00367</pub-id>
</mixed-citation>
</ref>
<ref id="B21">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Enfield</surname> <given-names>D. B.</given-names></name>
</person-group> (<year>2001</year>). 
<article-title>Evolution and historical perspective of the 1997&#x2013;1998 El Ni&#xf1;o&#x2013;Southern Oscillation event</article-title>. <source>Bull. Mar. Sci.</source> <volume>69</volume>, <fpage>7</fpage>&#x2013;<lpage>25</lpage>.
</mixed-citation>
</ref>
<ref id="B22">
<mixed-citation publication-type="web">
<person-group person-group-type="author"><collab>ERAS</collab>
</person-group>. (<year>2025</year>). 
<article-title>ERA Explorer Climate Data Portal</article-title>. <source>Copernicus Climate Change Service</source>. Available online at:&#xa0;<uri xlink:href="https://era-explorer.climate.copernicus.eu/?lat=-2.51&amp;lng=-80.07&amp;plot=3">https://era-explorer.climate.copernicus.eu/?lat=-2.51&amp;lng=-80.07&amp;plot=3</uri>
</mixed-citation>
</ref>
<ref id="B23">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Espinoza-Morriber&#xf3;n</surname> <given-names>D.</given-names></name>
<name><surname>Echevin</surname> <given-names>V.</given-names></name>
<name><surname>Colas</surname> <given-names>F.</given-names></name>
<name><surname>D&#xed;az</surname> <given-names>E.</given-names></name>
<name><surname>Tam</surname> <given-names>J.</given-names></name>
<name><surname>Anculle</surname> <given-names>T.</given-names></name>
<etal/>
</person-group>. (<year>2021</year>). 
<article-title>Diferencias entre los impactos en la costa Peruana de los eventos ENOS c&#xe1;lidos y El Ni&#xf1;o Costero 2017: vientos, afloramiento, productividad y anchoveta</article-title>. <source>Bol Inst Del Mar. Del PE.</source> <volume>36</volume>, <fpage>329</fpage>&#x2013;<lpage>348</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.53554/boletin.v36i2.34</pub-id>
</mixed-citation>
</ref>
<ref id="B24">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Fahrin</surname> <given-names>F.</given-names></name>
<name><surname>Gonzalez</surname> <given-names>A. O.</given-names></name>
<name><surname>Chrisler</surname> <given-names>B.</given-names></name>
<name><surname>Stachnik</surname> <given-names>J. P.</given-names></name>
</person-group> (<year>2024</year>). 
<article-title>The relationship between convectively coupled waves and the east pacific ITCZ</article-title>. <source>J. Climate</source> <volume>37</volume>, <fpage>2565</fpage>&#x2013;<lpage>2583</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1175/JCLI-D-23-0398.1</pub-id>
</mixed-citation>
</ref>
<ref id="B25">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Folland</surname> <given-names>C. K.</given-names></name>
<name><surname>Renwick</surname> <given-names>J. A.</given-names></name>
<name><surname>Salinger</surname> <given-names>M. J.</given-names></name>
<name><surname>Mullan</surname> <given-names>A. B.</given-names></name>
</person-group> (<year>2002</year>). 
<article-title>Relative influences of the interdecadal Pacific oscillation and ENSO on the South Pacific convergence zone</article-title>. <source>Geophys. Res. Lett.</source> <volume>29</volume>, <fpage>21</fpage>&#x2013;<lpage>21</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1029/2001GL014201</pub-id>
</mixed-citation>
</ref>
<ref id="B26">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Garc&#xe9;s-Vargas</surname> <given-names>J.</given-names></name>
<name><surname>Schneider</surname> <given-names>W.</given-names></name>
<name><surname>del R&#xed;o</surname> <given-names>R. A.</given-names></name>
<name><surname>Mart&#xed;nez</surname> <given-names>R.</given-names></name>
<name><surname>Zambrano</surname> <given-names>E.</given-names></name>
</person-group> (<year>2005</year>). 
<article-title>Inter-annual variability in the thermal structure of an oceanic time series station off Ecuado&#x2013;2003) associated with El Nino events</article-title>. <source>Dp. Sea. Res. Part I.: Oceanogr. Res. Pap.</source> <volume>52</volume>, <fpage>1789</fpage>&#x2013;<lpage>1805</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1016/j.dsr.2005.05.008</pub-id>
</mixed-citation>
</ref>
<ref id="B27">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Garreaud</surname> <given-names>R. D.</given-names></name>
</person-group> (<year>2018</year>). 
<article-title>A plausible atmospheric trigger for the 2017 coastal El Ni&#xf1;o</article-title>. <source>Int. J. Climatol</source> <volume>38</volume>, <fpage>e1296</fpage>&#x2013;<lpage>e1302</lpage>. Available online at:     <uri xlink:href="https://repositorio.uChile.cl/handle/2250/152026">https://repositorio.uChile.cl/handle/2250/152026</uri> (Accessed <date-in-citation content-type="access-date">June 15, 2025</date-in-citation>).
</mixed-citation>
</ref>
<ref id="B28">
<mixed-citation publication-type="book">
<person-group person-group-type="author">
<name><surname>Glantz</surname> <given-names>M. H.</given-names></name>
</person-group> (<year>2001</year>). <source>Once Burned, Twice Shy? Lessons Learned from the 1997&#x2013;1998 El Ni&#x2dc;no</source> (<publisher-loc>Tokyo-Japan</publisher-loc>: 
<publisher-name>The Unite Nations University</publisher-name>), <fpage>294</fpage>.
</mixed-citation>
</ref>
<ref id="B29">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Halpern</surname> <given-names>D.</given-names></name>
</person-group> (<year>1987</year>). 
<article-title>Observations of annual and El Ni&#xf1;o thermal and flow variations at 0&#xb0;, 110&#xb0;W and 0&#xb0;, 95&#xb0;W during 1980&#x2013;1985</article-title>. <source>J. Geophys. Res.</source> <volume>92</volume>, <fpage>8197</fpage>&#x2013;<lpage>8212</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1029/JC092iC08p08197</pub-id>
</mixed-citation>
</ref>
<ref id="B30">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Hansen</surname> <given-names>D. V.</given-names></name>
<name><surname>Poulain</surname> <given-names>P. M.</given-names></name>
</person-group> (<year>1995</year>). 
<article-title>Quality control and interpolations of WOCE-TOGA drifter data</article-title>. <source>J. Atmos. Ocean. Technol.</source> <volume>12</volume>, <fpage>900</fpage>&#x2013;<lpage>909</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1175/1520-0426(1995)012&lt;0900:QCAIOW&gt;2.0.CO;2</pub-id>
</mixed-citation>
</ref>
<ref id="B31">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Huang</surname> <given-names>Y.</given-names></name>
<name><surname>Shu</surname> <given-names>M.</given-names></name>
<name><surname>Liu</surname> <given-names>F.</given-names></name>
<name><surname>Liu</surname> <given-names>B.</given-names></name>
<name><surname>Huang</surname> <given-names>C.</given-names></name>
<name><surname>Wang</surname> <given-names>S.</given-names></name>
</person-group> (<year>2025</year>). 
<article-title>Nonlinear heterogeneity impact of El Ni&#xf1;o-Southern Oscillation on energy markets: A global perspective analysis</article-title>. <source>Energy</source> <volume>333</volume>, <fpage>137475</fpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1016/j.energy.2025.137475</pub-id>
</mixed-citation>
</ref>
<ref id="B32">
<mixed-citation publication-type="web">
<person-group person-group-type="author"><collab>IOC</collab>
</person-group> (<year>2018</year>). <source>Sixth international XBT science workshop, IOC project office for IODE</source>. Available online at: <uri xlink:href="http://www.iode.org/wr283">http://www.iode.org/wr283</uri> (Accessed <date-in-citation content-type="access-date">June 24, 2022</date-in-citation>).
</mixed-citation>
</ref>
<ref id="B33">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Islam</surname> <given-names>F. S.</given-names></name>
</person-group> (<year>2025</year>). 
<article-title>Assessment of the global climatic impacts due to el nino and la nina events</article-title>. <source>J. Global Ecol. Environ.</source> <volume>21</volume>, <fpage>1</fpage>&#x2013;<lpage>26</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.56557/jogee/2025/v21i39333</pub-id>
</mixed-citation>
</ref>
<ref id="B34">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Johnson</surname> <given-names>G. C.</given-names></name>
<name><surname>Lyman</surname> <given-names>J. M.</given-names></name>
</person-group> (<year>2022</year>). 
<article-title>GOSML: A global ocean surface mixed layer statistical monthly climatology: Means, percentiles, skewness, and kurtosis</article-title>. <source>J. Geophys. Res.: Ocean.</source> <volume>127</volume>, <fpage>e2021JC018219</fpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1029/2021JC018219</pub-id>
</mixed-citation>
</ref>
<ref id="B35">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Kent</surname> <given-names>E. C.</given-names></name>
<name><surname>Fangohr</surname> <given-names>S.</given-names></name>
<name><surname>Berry</surname> <given-names>D. I.</given-names></name>
</person-group> (<year>2012</year>). 
<article-title>A comparative assessment of monthly mean wind speed products over the global ocean</article-title>. <source>Int. J. Climatol.</source> <volume>33</volume> (<issue>11</issue>). doi:&#xa0;<pub-id pub-id-type="doi">10.1002/joc.3606</pub-id>
</mixed-citation>
</ref>
<ref id="B36">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Kent</surname> <given-names>C.</given-names></name>
<name><surname>Scaife</surname> <given-names>A. A.</given-names></name>
<name><surname>Seviour</surname> <given-names>W. J.</given-names></name>
<name><surname>Dunstone</surname> <given-names>N.</given-names></name>
<name><surname>Smith</surname> <given-names>D.</given-names></name>
<name><surname>Ineson</surname> <given-names>S.</given-names></name>
</person-group> (<year>2025</year>). 
<article-title>Transition of El Ni&#xf1;o to La Ni&#xf1;a can be driven by regional perturbations a year ahead</article-title>. <source>Environ. Res. Lett.</source> <volume>20</volume>, <fpage>064014</fpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1088/1748-9326/add35d</pub-id>
</mixed-citation>
</ref>
<ref id="B37">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Kusuma</surname> <given-names>D. W.</given-names></name>
<name><surname>Al Ma&#x2019;rufi</surname> <given-names>I.</given-names></name>
<name><surname>Muslim</surname> <given-names>M. A.</given-names></name>
</person-group> (<year>2018</year>). 
<article-title>Spatial interpolation of sea surface temperature using kriging and inverse distance weighting methods in Java Sea, Indonesia</article-title>. <source>IOP. Conf. Ser.: Earth Environ. Sci.</source> <volume>116</volume>, <elocation-id>12031</elocation-id>. doi:&#xa0;<pub-id pub-id-type="doi">10.1088/1755-1315/116/1/012031</pub-id>
</mixed-citation>
</ref>
<ref id="B38">
<mixed-citation publication-type="book">
<person-group person-group-type="author">
<name><surname>Le</surname> <given-names>N. D.</given-names></name>
<name><surname>Zidek</surname> <given-names>J. V.</given-names></name>
</person-group> (<year>2006</year>). <source>Statistical Analysis of Environmental Space-Time Processes</source> (<publisher-loc>Cham, Switzerland, at Gewerbestrasse 11</publisher-loc>: 
<publisher-name>Springer</publisher-name>). doi:&#xa0;<pub-id pub-id-type="doi">10.1007/0-387-31143-8</pub-id>
</mixed-citation>
</ref>
<ref id="B39">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Li</surname> <given-names>X.</given-names></name>
<name><surname>Hu</surname> <given-names>Z.-Z.</given-names></name>
<name><surname>McPhaden</surname> <given-names>M. J.</given-names></name>
<name><surname>Zhu</surname> <given-names>C.</given-names></name>
<name><surname>Liu</surname> <given-names>Y.</given-names></name>
</person-group> (<year>2023</year>). 
<article-title>Triple-Dip la ni&#xf1;as in 1998&#x2013;2001 and 2020&#x2013;2023: impact of mean state changes</article-title>. <source>J. Geophys. Res.: Atmos.</source> <volume>128</volume>, <elocation-id>e2023JD038843</elocation-id>. doi:&#xa0;<pub-id pub-id-type="doi">10.1029/2023JD038843</pub-id>
</mixed-citation>
</ref>
<ref id="B40">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Li</surname> <given-names>Z. L.</given-names></name>
<name><surname>Tang</surname> <given-names>B. H.</given-names></name>
<name><surname>Wu</surname> <given-names>H.</given-names></name>
<name><surname>Ren</surname> <given-names>H.</given-names></name>
<name><surname>Yan</surname> <given-names>G.</given-names></name>
<name><surname>Wan</surname> <given-names>Z.</given-names></name>
<etal/>
</person-group>. (<year>2013</year>). 
<article-title>Satellite-derived land surface temperature: Current status and perspectives</article-title>. <source>Remote Sens. Environ.</source> <volume>131</volume>, <fpage>14</fpage>&#x2013;<lpage>37</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1016/j.rse.2012.12.008</pub-id>
</mixed-citation>
</ref>
<ref id="B41">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Li</surname> <given-names>X.</given-names></name>
<name><surname>Yu</surname> <given-names>J.-Y.</given-names></name>
<name><surname>Ding</surname> <given-names>R.</given-names></name>
</person-group> (<year>2024</year>). 
<article-title>El Ni&#xf1;o-La Ni&#xf1;a asymmetries in the changes of ENSO complexities and dynamics since 1990</article-title>. <source>Geophys. Res. Lett.</source> <volume>51</volume>, <fpage>e2023GL106395</fpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1029/2023GL106395</pub-id>
</mixed-citation>
</ref>
<ref id="B42">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>L&#xfc;bbecke</surname> <given-names>J. F.</given-names></name>
<name><surname>Rudloff</surname> <given-names>D.</given-names></name>
<name><surname>Stramma</surname> <given-names>L.</given-names></name>
</person-group> (<year>2019</year>). 
<article-title>Stand-alone eastern Pacific Coastal Warming events</article-title>. <source>Geophys. Res. Lett.</source> <volume>46</volume>, <fpage>12360</fpage>&#x2013;<lpage>12367</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1029/2019GL084479</pub-id>
</mixed-citation>
</ref>
<ref id="B43">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Mangum</surname> <given-names>L. J.</given-names></name>
<name><surname>Hayes</surname> <given-names>S. P.</given-names></name>
<name><surname>Toole</surname> <given-names>J. M.</given-names></name>
</person-group> (<year>1986</year>). 
<article-title>Eastern Pacific Ocean circulation near the onset of the 1982&#x2013;1983 El Ni&#xf1;o</article-title>. <source>J. Geophys. Res.</source> <volume>91</volume>, <fpage>8428</fpage>&#x2013;<lpage>8436</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1029/JC091iC07p08428</pub-id>
</mixed-citation>
</ref>
<ref id="B44">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Margaritis</surname> <given-names>G.</given-names></name>
<name><surname>Kent</surname> <given-names>E. C.</given-names></name>
<name><surname>Foster</surname> <given-names>G. L.</given-names></name>
</person-group> (<year>2025</year>). 
<article-title>Intercomparison of satellite-derived SST with logger data in the Caribbean&#x2014;Implications for coral reef monitoring</article-title>. <source>PloS Climate</source> <volume>4</volume>, <fpage>e0000480</fpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1371/journal.pclm.0000480</pub-id>
</mixed-citation>
</ref>
<ref id="B45">
<mixed-citation publication-type="book">
<person-group person-group-type="author">
<name><surname>Marsh</surname> <given-names>T.</given-names></name>
</person-group> (<year>2011</year>). <source>Atmospheric Moisture, Precipitation, and Weather Systems</source> (<publisher-loc>Cambridge, UK</publisher-loc>: 
<publisher-name>Cambridge University Press</publisher-name>).
</mixed-citation>
</ref>
<ref id="B46">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>McPhaden</surname> <given-names>M. J.</given-names></name>
<name><surname>Zebiak</surname> <given-names>S. E.</given-names></name>
<name><surname>Glantz</surname> <given-names>M. H.</given-names></name>
</person-group> (<year>2006</year>). 
<article-title>ENSO as an integrating concept in Earth science</article-title>. <source>Science</source> <volume>314</volume>, <fpage>1740</fpage>&#x2013;<lpage>1745</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1126/science.1132588</pub-id>, PMID: <pub-id pub-id-type="pmid">17170296</pub-id>
</mixed-citation>
</ref>
<ref id="B47">
<mixed-citation publication-type="web">
<person-group person-group-type="author"><collab>MEI</collab>
</person-group> (<year>2024</year>). <source>MEI.v2: NOAA Physical Sciences Laboratory</source>.
</mixed-citation>
</ref>
<ref id="B48">
<mixed-citation publication-type="book">
<person-group person-group-type="author"><collab>MetOffice</collab>
</person-group> (<year>2024</year>). <source>El Ni&#xf1;o Southern Oscillation (ENSO) region sea surface temperature forecasts</source> (<publisher-loc>Exeter, Devon, UK</publisher-loc>: 
<publisher-name>Met Office</publisher-name>).
</mixed-citation>
</ref>
<ref id="B49">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Michaud</surname> <given-names>F.</given-names></name>
<name><surname>Proust</surname> <given-names>J. N.</given-names></name>
<name><surname>Collot</surname> <given-names>J. Y.</given-names></name>
<name><surname>Lebrun</surname> <given-names>J. F.</given-names></name>
<name><surname>Witt</surname> <given-names>C.</given-names></name>
<name><surname>Ratzov</surname> <given-names>G.</given-names></name>
<etal/>
</person-group>. (<year>2015</year>). 
<article-title>Quaternary sedimentation and active faulting along the Ecuadorian shelf: preliminary results of the ATACAMES Cruise, (2012)</article-title>. <source>Mar. Geophys. Res.</source> <volume>36</volume>, <fpage>81</fpage>&#x2013;<lpage>98</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1007/s11001-014-9231-y</pub-id>
</mixed-citation>
</ref>
<ref id="B50">
<mixed-citation publication-type="book">
<person-group person-group-type="author">
<name><surname>Michaud</surname> <given-names>F.</given-names></name>
<name><surname>Proust</surname> <given-names>J. N.</given-names></name>
<name><surname>Collot</surname> <given-names>J. Y.</given-names></name>
<name><surname>Lebrun</surname> <given-names>J. F.</given-names></name>
<name><surname>Witt</surname> <given-names>C.</given-names></name>
<name><surname>Ratzov</surname> <given-names>G.</given-names></name>
<etal/>
</person-group>. (<year>2024</year>). <source>SUPER-MOUV &#x2013; Cruise Report: Sub-seafloor effects of the Pedernales Earthquake rupture/Ecuador</source> (<publisher-loc>France</publisher-loc>: 
<publisher-name>R&#xe9;sultats pr&#xe9;liminaires, CNRS&#x2013;IRD&#x2013;Universit&#xe9; C&#xf4;te d&#x2019;Azur</publisher-name>), <fpage>725</fpage>.
</mixed-citation>
</ref>
<ref id="B51">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Mitsopoulos</surname> <given-names>P.</given-names></name>
<name><surname>Pe&#xf1;a</surname> <given-names>M.</given-names></name>
</person-group> (<year>2023</year>). 
<article-title>Characterizing coastal wind speed and significant wave height using satellite altimetry and buoy data</article-title>. <source>Remote Sens.</source> <volume>15</volume>, <elocation-id>987</elocation-id>. doi:&#xa0;<pub-id pub-id-type="doi">10.3390/rs15040987</pub-id>
</mixed-citation>
</ref>
<ref id="B52">
<mixed-citation publication-type="web">
<person-group person-group-type="author"><collab>NOAA</collab>
</person-group> (<year>2023</year>). Available online at: <uri xlink:href="https://www.cpc.ncep.noaa.gov/">https://www.cpc.ncep.noaa.gov/</uri> (Accessed <date-in-citation content-type="access-date">December 20, 2024</date-in-citation>).
</mixed-citation>
</ref>
<ref id="B53">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>O&#x2019;Dea</surname> <given-names>A.</given-names></name>
<name><surname>Sellers</surname> <given-names>A. J.</given-names></name>
<name><surname>P&#xe9;rez-Medina</surname> <given-names>C.</given-names></name>
<name><surname>Pardo D&#xed;az</surname> <given-names>J.</given-names></name>
<name><surname>Guzm&#xe1;n Bloise</surname> <given-names>A.</given-names></name>
<name><surname>P&#xf6;hlker</surname> <given-names>C.</given-names></name>
<etal/>
</person-group>. (<year>2025</year>). 
<article-title>Unprecedented suppression of Panama&#x2019;s Pacific upwelling in 2025</article-title>. <source>Proc. Natl. Acad. Sci. U.S.A.</source> <volume>122</volume>, <fpage>e2512056122</fpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1073/pnas.2512056122</pub-id>, PMID: <pub-id pub-id-type="pmid">40892925</pub-id>
</mixed-citation>
</ref>
<ref id="B54">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Ord&#xf3;&#xf1;ez-Z&#xfa;&#xf1;iga</surname> <given-names>S. A.</given-names></name>
<name><surname>Correa-Ram&#xed;rez</surname> <given-names>M.</given-names></name>
<name><surname>Ricaurte-Villota</surname> <given-names>C.</given-names></name>
<name><surname>Bastidas-Salamanca</surname> <given-names>M.</given-names></name>
</person-group> (<year>2021</year>). 
<article-title>The Panama Low-Level Jet: extension, annual cycle and modes of variation</article-title>. <source>Lat. Am. J. Aquat. Res.</source> <volume>49</volume>, <fpage>750</fpage>&#x2013;<lpage>762</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.3856/vol49-issue5-fulltext-2591</pub-id>
</mixed-citation>
</ref>
<ref id="B55">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Ormaza-Gonz&#xe1;lez</surname> <given-names>F. I.</given-names></name>
</person-group> (<year>2016</year>). 
<article-title>Oceanographic events of high and low frequency in the eastern equatorial Pacific Ocean. Evaluation of its monitoring and prediction</article-title>. <source>Rev. Int. Invest y. Doc. (RIID)</source> <volume>1</volume>, <fpage>2445</fpage>&#x2013;<lpage>1711</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.19239/riidv1n3p1</pub-id>
</mixed-citation>
</ref>
<ref id="B56">
<mixed-citation publication-type="book">
<person-group person-group-type="author">
<name><surname>Ormaza-Gonz&#xe1;lez</surname> <given-names>F. I.</given-names></name>
</person-group> (<year>2024</year>). <source>The South Pacific Anticyclone and its impact on Ecuador: A beneficial natural phenomenon for the region. The South Pacific Anticyclone and its impact on Ecuador: A beneficial natural phenomenon for the region</source> (<publisher-loc>Guayaquil, Ecuador</publisher-loc>: 
<publisher-name>FIMCM</publisher-name>).
</mixed-citation>
</ref>
<ref id="B57">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Ormaza-Gonz&#xe1;lez</surname> <given-names>F. I.</given-names></name>
<name><surname>Cede&#xf1;o</surname> <given-names>J.</given-names></name>
</person-group> (<year>2017</year>). 
<article-title>Coastal El Ni&#xf1;o 2017 or simply: the carnival coastal warming event</article-title>? <source>MOJ. Ecol. Environ. Sci.</source> <volume>2</volume>, <fpage>54</fpage>&#x2013;<lpage>57</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.15406/mojes.2017.02.0005400054</pub-id>
</mixed-citation>
</ref>
<ref id="B58">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Ormaza-Gonz&#xe1;lez</surname> <given-names>F. I.</given-names></name>
<name><surname>Espinoza-Celi</surname> <given-names>Mar&#xed;aE.</given-names></name>
<name><surname>Roa-L&#xf3;pez</surname> <given-names>H. M.</given-names></name>
</person-group> (<year>2022</year>). 
<article-title>Did Schwabe cycles 19&#x2013;24 influence the ENSO events, PDO, and AMO indexes in the Pacific and Atlantic Oceans</article-title>? <source>Global Planet. Change</source> <volume>217</volume>, <elocation-id>103928</elocation-id>. doi:&#xa0;<pub-id pub-id-type="doi">10.1016/j.gloplacha.2022.103928</pub-id>
</mixed-citation>
</ref>
<ref id="B59">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Parks</surname> <given-names>J.</given-names></name>
<name><surname>Bringas</surname> <given-names>F.</given-names></name>
<name><surname>Cowley</surname> <given-names>R.</given-names></name>
<name><surname>Hanstein</surname> <given-names>C.</given-names></name>
<name><surname>Krummel</surname> <given-names>L.</given-names></name>
<name><surname>Sprintall</surname> <given-names>J.</given-names></name>
<etal/>
</person-group>. (<year>2022</year>). 
<article-title>XBT operational best practices for quality assurance</article-title>. <source>Front. Mar. Sci.</source> <volume>9</volume>. doi:&#xa0;<pub-id pub-id-type="doi">10.3389/fmars.2022.991760</pub-id>
</mixed-citation>
</ref>
<ref id="B60">
<mixed-citation publication-type="book">
<person-group person-group-type="author"><collab>PDO</collab>
</person-group> (<year>2024</year>). <source>Pacific Decadal Oscillation. Pacific Decadal Oscillation (PDO)</source> (<publisher-loc>Asheville, NC (Headquarters), USA</publisher-loc>: 
<publisher-name>National Centers for Environmental Information (NCEI</publisher-name>).
</mixed-citation>
</ref>
<ref id="B61">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Peng</surname> <given-names>Q.</given-names></name>
<name><surname>Xie</surname> <given-names>S. P.</given-names></name>
<name><surname>Passalacqua</surname> <given-names>G. A.</given-names></name>
<name><surname>Miyamoto</surname> <given-names>A.</given-names></name>
<name><surname>Deser</surname> <given-names>C.</given-names></name>
</person-group> (<year>2024</year>). 
<article-title>The 2023 extreme coastal El Ni&#xf1;o: Atmospheric and air-sea coupling mechanisms</article-title>. <source>Sci. Adv.</source> <volume>10</volume>, <elocation-id>eadk8646</elocation-id>. doi:&#xa0;<pub-id pub-id-type="doi">10.1126/sciadv.adk8646</pub-id>, PMID: <pub-id pub-id-type="pmid">38517959</pub-id>
</mixed-citation>
</ref>
<ref id="B62">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Peters</surname> <given-names>B. D.</given-names></name>
<name><surname>Jenkins</surname> <given-names>W. J.</given-names></name>
<name><surname>Swift</surname> <given-names>J. H.</given-names></name>
<name><surname>German</surname> <given-names>C. R.</given-names></name>
<name><surname>Moffett</surname> <given-names>J. W.</given-names></name>
<name><surname>Cutter</surname> <given-names>G. A.</given-names></name>
<etal/>
</person-group>. (<year>2018</year>). 
<article-title>Water mass analysis of the 2013 US GEOTRACES eastern Pacific zonal transect (GP16)</article-title>. <source>Mar. Chem.</source> <volume>201</volume>, <fpage>6</fpage>&#x2013;<lpage>19</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1016/j.marchem.2017.09.007</pub-id>
</mixed-citation>
</ref>
<ref id="B63">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Ram&#xed;rez</surname> <given-names>I. J.</given-names></name>
<name><surname>Briones</surname> <given-names>F.</given-names></name>
</person-group> (<year>2017</year>). 
<article-title>Understanding the El ni&#xf1;o costero of 2017: the definition problem and challenges of climate forecasting and disaster responses</article-title>. <source>Int. J. Disast. Risk Sci.</source> <volume>8</volume>, <fpage>489</fpage>&#x2013;<lpage>492</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1007/s13753-017-0151-8</pub-id>
</mixed-citation>
</ref>
<ref id="B64">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Rollenbeck</surname> <given-names>R.</given-names></name>
<name><surname>Orellana-Alvear</surname> <given-names>J.</given-names></name>
<name><surname>Bendix</surname> <given-names>J.</given-names></name>
<name><surname>Rodriguez</surname> <given-names>R.</given-names></name>
<name><surname>Pucha-Cofrep</surname> <given-names>F.</given-names></name>
<name><surname>Guallpa</surname> <given-names>M.</given-names></name>
<etal/>
</person-group>. (<year>2022</year>). 
<article-title>The coastal el ni&#xf1;o event of 2017 in Ecuador and Peru: A weather radar analysis</article-title>. <source>Remote Sens.</source> <volume>14</volume>, <elocation-id>824</elocation-id>. doi:&#xa0;<pub-id pub-id-type="doi">10.3390/rs14040824</pub-id>
</mixed-citation>
</ref>
<ref id="B65">
<mixed-citation publication-type="book">
<person-group person-group-type="author">
<name><surname>Saltos-Aguilar</surname> <given-names>D.</given-names></name>
<name><surname>Ormaza-Gonz&#xe1;lez</surname> <given-names>F. I.</given-names></name>
</person-group> (<year>2023</year>). <source>Condiciones oceanogr&#xe1;ficas durante septiembre 2023</source> (
<publisher-name>C&#xe1;mara Nacional de Pesquer&#xed;as del Ecuador</publisher-name>). Available online at: <uri xlink:href="https://camaradepesqueria.ec/condiciones-oceanograficas-durante-septiembre-2023/">https://camaradepesqueria.ec/condiciones-oceanograficas-durante-septiembre-2023/</uri> (Accessed <date-in-citation content-type="access-date">March, 20, 2025</date-in-citation>).
</mixed-citation>
</ref>
<ref id="B66">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Takahashi</surname> <given-names>K.</given-names></name>
<name><surname>Mart&#xed;nez</surname> <given-names>A. G.</given-names></name>
</person-group> (<year>2019</year>). 
<article-title>The very strong coastal El Ni&#xf1;o in 1925 in the far-eastern Pacific</article-title>. <source>Climate Dyn.</source> <volume>52</volume>, <fpage>7389</fpage>&#x2013;<lpage>7415</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1007/s00382-017-3702-1</pub-id>
</mixed-citation>
</ref>
<ref id="B67">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Tan</surname> <given-names>W.</given-names></name>
<name><surname>Hu</surname> <given-names>Z. Z.</given-names></name>
<name><surname>McPhaden</surname> <given-names>M. J.</given-names></name>
<name><surname>Zhu</surname> <given-names>C.</given-names></name>
<name><surname>Li</surname> <given-names>X.</given-names></name>
<name><surname>Liu</surname> <given-names>Y.</given-names></name>
</person-group> (<year>2024</year>). 
<article-title>On the divergent evolution of ENSO after the coastal El Ni&#xf1;os in 2017 and 2023</article-title>. <source>Geophys. Res. Lett.</source> <volume>51</volume>, <fpage>e2024GL108198</fpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1029/2024GL108198</pub-id>
</mixed-citation>
</ref>
<ref id="B68">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Thielen</surname> <given-names>D. R.</given-names></name>
<name><surname>Ramoni-Perazzi</surname> <given-names>P.</given-names></name>
<name><surname>Zamora-Ledezma</surname> <given-names>E.</given-names></name>
<name><surname>Puche</surname> <given-names>M. L.</given-names></name>
<name><surname>Marquez</surname> <given-names>M.</given-names></name>
<name><surname>Quintero</surname> <given-names>J. I.</given-names></name>
<etal/>
</person-group>. (<year>2023</year>). 
<article-title>Effect of extreme El Ni&#xf1;o events on the precipitation of Ecuador</article-title>. <source>Nat. Hazard. Earth Syst. Sci.</source> <volume>23</volume>, <fpage>1507</fpage>&#x2013;<lpage>1527</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.5194/nhess-23-1507-2023</pub-id>
</mixed-citation>
</ref>
<ref id="B69">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Torres-Godoy</surname> <given-names>M.</given-names></name>
<name><surname>Pizarro</surname> <given-names>O.</given-names></name>
<name><surname>Dewitte</surname> <given-names>B.</given-names></name>
<name><surname>Oerder</surname> <given-names>V.</given-names></name>
</person-group> (<year>2025</year>). 
<article-title>ENSO-driven variability of deep ocean circulation in the southeast pacific</article-title>. <source>EGUsphere</source> <volume>2025</volume>, <fpage>1</fpage>&#x2013;<lpage>40</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.5194/egusphere-2025-1311</pub-id>
</mixed-citation>
</ref>
<ref id="B70">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Trenberth</surname> <given-names>K. E.</given-names></name>
</person-group> (<year>1997</year>). 
<article-title>The definition of el nino</article-title>. <source>Bull. Am. Meteorol. Soc.</source> <volume>78</volume>, <fpage>2771</fpage>&#x2013;<lpage>2778</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1175/1520-0477(1997)078&lt;2771:TDOENO&gt;2.0.CO;2</pub-id>
</mixed-citation>
</ref>
<ref id="B71">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Vialard</surname> <given-names>J.</given-names></name>
<name><surname>Jin</surname> <given-names>F. F.</given-names></name>
<name><surname>Mcphaden</surname> <given-names>M. J.</given-names></name>
<name><surname>Fedorov</surname> <given-names>A.</given-names></name>
<name><surname>Cai</surname> <given-names>W.</given-names></name>
<name><surname>An</surname> <given-names>S. I.</given-names></name>
<etal/>
</person-group>. (<year>2025</year>). 
<article-title>The El Ni&#xf1;o Southern Oscillation (ENSO) recharge oscillator conceptual model: Achievements and future prospects</article-title>. <source>Rev. Geophys.</source> <volume>63</volume>, <fpage>e2024RG000843</fpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1029/2024RG000843</pub-id>
</mixed-citation>
</ref>
<ref id="B72">
<mixed-citation publication-type="book">
<person-group person-group-type="author">
<name><surname>Wang</surname> <given-names>C.</given-names></name>
<name><surname>Deser</surname> <given-names>C.</given-names></name>
<name><surname>Yu</surname> <given-names>J. Y.</given-names></name>
<name><surname>DiNezio</surname> <given-names>P.</given-names></name>
<name><surname>Clement</surname> <given-names>A.</given-names></name>
</person-group> (<year>2017</year>). &#x201c;
<article-title>El Ni&#xf1;o and Southern Oscillation (ENSO): A Review</article-title>,&#x201d; in <source>Coral Reefs of the Eastern Tropical Pacific. Coral Reefs of the World</source>, vol. <volume>8</volume> . Eds. 
<person-group person-group-type="editor">
<name><surname>Glynn</surname> <given-names>P.</given-names></name>
<name><surname>Manzello</surname> <given-names>D.</given-names></name>
<name><surname>Enochs</surname> <given-names>I.</given-names></name>
</person-group> (
<publisher-name>Springer</publisher-name>, <publisher-loc>Dordrecht</publisher-loc>). doi:&#xa0;<pub-id pub-id-type="doi">10.1007/978-94-017-7499-4_4</pub-id>
</mixed-citation>
</ref>
<ref id="B73">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Wang</surname> <given-names>C.</given-names></name>
<name><surname>Fiedler</surname> <given-names>P. C.</given-names></name>
</person-group> (<year>2006</year>). 
<article-title>ENSO variability and the eastern tropical Pacific: A review</article-title>. <source>Prog. Oceanogr.</source> <volume>69</volume>, <fpage>239</fpage>&#x2013;<lpage>266</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1016/j.pocean.2006.03.004</pub-id>
</mixed-citation>
</ref>
<ref id="B74">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Wang</surname> <given-names>J.</given-names></name>
<name><surname>Luo</surname> <given-names>H.</given-names></name>
<name><surname>Yu</surname> <given-names>L.</given-names></name>
<name><surname>Li</surname> <given-names>X.</given-names></name>
<name><surname>Holland</surname> <given-names>P. R.</given-names></name>
<name><surname>Yang</surname> <given-names>Q.</given-names></name>
</person-group> (<year>2023</year>). 
<article-title>The impacts of combined SAM and ENSO on seasonal Antarctic sea ice changes</article-title>. <source>Journal of Climate</source> <volume>36</volume> (<issue>11</issue>), <fpage>3553</fpage>&#x2013;<lpage>3569</lpage>.
</mixed-citation>
</ref>
<ref id="B75">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Webb</surname> <given-names>D. J.</given-names></name>
</person-group> (<year>2018</year>). 
<article-title>On the role of the North Equatorial Counter Current during a strong El Ni&#xf1;o</article-title>. <source>Ocean. Sci.</source> <volume>14</volume>, <fpage>633</fpage>&#x2013;<lpage>660</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.5194/os-14-633-2018</pub-id>
</mixed-citation>
</ref>
<ref id="B76">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Wyrtki</surname> <given-names>K.</given-names></name>
</person-group> (<year>1973</year>). 
<article-title>Teleconnections in the equatorial pacific ocean</article-title>. <source>Science</source> <volume>180</volume>, <fpage>66</fpage>&#x2013;<lpage>68</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1126/science.180.4081.66</pub-id>, PMID: <pub-id pub-id-type="pmid">17757976</pub-id>
</mixed-citation>
</ref>
<ref id="B77">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Wyrtki</surname> <given-names>K.</given-names></name>
</person-group> (<year>1974</year>). 
<article-title>Equatorial currents in the pacific 1950 to 1970 and their relation to the trade winds, J. Phys</article-title>. <source>Oceanogr</source> <volume>4</volume>, <fpage>372</fpage>&#x2013;<lpage>380</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1175/1520-0485(1974)004&lt;0372:ECITPT&gt;2.0.CO;2</pub-id>
</mixed-citation>
</ref>
<ref id="B78">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Yglesias-Gonz&#xe1;lez</surname> <given-names>M.</given-names></name>
<name><surname>Vald&#xe9;s-Vel&#xe1;squez</surname> <given-names>A.</given-names></name>
<name><surname>Hartinger</surname> <given-names>S. M.</given-names></name>
<name><surname>Takahashi</surname> <given-names>K.</given-names></name>
<name><surname>Salvatierra</surname> <given-names>G.</given-names></name>
<name><surname>Velarde</surname> <given-names>R.</given-names></name>
<etal/>
</person-group>. (<year>2023</year>). 
<article-title>Reflections on the impact and response to the Peruvian 2017 Coastal El Ni&#xf1;o event: Looking to the past to prepare for the future</article-title>. <source>PloS One</source> <volume>18</volume>, <fpage>e0290767</fpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1371/journal.pone.0290767</pub-id>, PMID: <pub-id pub-id-type="pmid">37751405</pub-id>
</mixed-citation>
</ref>
<ref id="B79">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Zhang</surname> <given-names>R.</given-names></name>
<name><surname>Levitus</surname> <given-names>S.</given-names></name>
</person-group> (<year>1997</year>). 
<article-title>Structure and Cycle of Decadal Variability of Upper-Ocean Temperature in the North Pacific</article-title>. <source>Journal of Climate</source> <volume>10</volume>, <fpage>710</fpage>&#x2013;<lpage>727</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1175/1520-0442(1997)010&lt;0710:SACODV&gt;2.0.CO;2</pub-id>
</mixed-citation>
</ref>
<ref id="B80">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Zhang</surname> <given-names>S.</given-names></name>
<name><surname>Fan</surname> <given-names>H.</given-names></name>
<name><surname>Hu</surname> <given-names>X.</given-names></name>
<name><surname>Lin</surname> <given-names>S.</given-names></name>
</person-group> (<year>2024</year>). 
<article-title>Unprecedented cross-equatorial southerly wind anomalies during the 2020&#x2013;2023 triple-dip La Ni&#xf1;a: Impacts and mechanisms</article-title>. <source>Atmos. Res.</source> <volume>304</volume>, <elocation-id>107412</elocation-id>. doi:&#xa0;<pub-id pub-id-type="doi">10.1016/j.atmosres.2024.107412</pub-id>
</mixed-citation>
</ref>
<ref id="B81">
<mixed-citation publication-type="journal">
<person-group person-group-type="author">
<name><surname>Zhao</surname> <given-names>J.</given-names></name>
<name><surname>Li</surname> <given-names>Y.</given-names></name>
<name><surname>Wang</surname> <given-names>F.</given-names></name>
</person-group> (<year>2013</year>). 
<article-title>Dynamical responses of the west Pacific North Equatorial Countercurrent (NECC) system to El Ni&#xf1;o events</article-title>. <source>J. Geophys. Res. Ocean.</source> <volume>118</volume>, <fpage>2828</fpage>&#x2013;<lpage>2844</lpage>. doi:&#xa0;<pub-id pub-id-type="doi">10.1002/jgrc.20196</pub-id>
</mixed-citation>
</ref>
</ref-list>
<fn-group>
<fn id="n1" fn-type="custom" custom-type="edited-by">
<p>Edited by: <ext-link ext-link-type="uri" xlink:href="https://loop.frontiersin.org/people/616471">Juan Jose Munoz-Perez</ext-link>, University of C&#xe1;diz, Spain</p></fn>
<fn id="n2" fn-type="custom" custom-type="reviewed-by">
<p>Reviewed by: <ext-link ext-link-type="uri" xlink:href="https://loop.frontiersin.org/people/1462395">Edgardo Diaz-Ferguson</ext-link>, Estaci&#xf3;n Cient&#xed;fica Coiba AIP, Panama</p>
<p><ext-link ext-link-type="uri" xlink:href="https://loop.frontiersin.org/people/1854950">Yuting Feng</ext-link>, Independent researcher, St Petersburg, United States</p>
<p><ext-link ext-link-type="uri" xlink:href="https://loop.frontiersin.org/people/3287740">Gustavo Cardenas</ext-link>, University of Panama, Panama</p></fn>
</fn-group>
</back>
</article>