<?xml version="1.0" encoding="UTF-8" standalone="no"?>
<!DOCTYPE article PUBLIC "-//NLM//DTD Journal Publishing DTD v2.3 20070202//EN" "journalpublishing.dtd">
<article xml:lang="EN" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" article-type="research-article">
<front>
<journal-meta>
<journal-id journal-id-type="publisher-id">Front. For. Glob. Change</journal-id>
<journal-title>Frontiers in Forests and Global Change</journal-title>
<abbrev-journal-title abbrev-type="pubmed">Front. For. Glob. Change</abbrev-journal-title>
<issn pub-type="epub">2624-893X</issn>
<publisher>
<publisher-name>Frontiers Media S.A.</publisher-name>
</publisher>
</journal-meta>
<article-meta>
<article-id pub-id-type="doi">10.3389/ffgc.2024.1464140</article-id>
<article-categories>
<subj-group subj-group-type="heading">
<subject>Forests and Global Change</subject>
<subj-group>
<subject>Original Research</subject>
</subj-group>
</subj-group>
</article-categories>
<title-group>
<article-title>Impact of winch-assisted logging machinery on soil disturbance in the mountainous forests of Western Carpathians</article-title>
</title-group>
<contrib-group>
<contrib contrib-type="author">
<name><surname>Allman</surname> <given-names>Michal</given-names></name>
<xref ref-type="aff" rid="aff1"><sup>1</sup></xref>
<uri xlink:href="http://loop.frontiersin.org/people/2837705/overview"/>
<role content-type="https://credit.niso.org/contributor-roles/writing-original-draft/"/>
<role content-type="https://credit.niso.org/contributor-roles/writing-review-editing/"/>
<role content-type="https://credit.niso.org/contributor-roles/conceptualization/"/>
<role content-type="https://credit.niso.org/contributor-roles/data-curation/"/>
<role content-type="https://credit.niso.org/contributor-roles/methodology/"/>
<role content-type="https://credit.niso.org/contributor-roles/software/"/>
<role content-type="https://credit.niso.org/contributor-roles/visualization/"/>
</contrib>
<contrib contrib-type="author">
<name><surname>Dud&#x000E1;kov&#x000E1;</surname> <given-names>Zuzana</given-names></name>
<xref ref-type="aff" rid="aff1"><sup>1</sup></xref>
<xref ref-type="author-notes" rid="fn001"><sup>&#x02020;</sup></xref>
<role content-type="https://credit.niso.org/contributor-roles/writing-original-draft/"/>
<role content-type="https://credit.niso.org/contributor-roles/writing-review-editing/"/>
<role content-type="https://credit.niso.org/contributor-roles/conceptualization/"/>
<role content-type="https://credit.niso.org/contributor-roles/formal-analysis/"/>
<role content-type="https://credit.niso.org/contributor-roles/methodology/"/>
<role content-type="https://credit.niso.org/contributor-roles/visualization/"/>
</contrib>
<contrib contrib-type="author" corresp="yes">
<name><surname>Duchan</surname> <given-names>Martin</given-names></name>
<xref ref-type="aff" rid="aff1"><sup>1</sup></xref>
<xref ref-type="corresp" rid="c001"><sup>&#x0002A;</sup></xref>
<uri xlink:href="http://loop.frontiersin.org/people/2789191/overview"/>
<role content-type="https://credit.niso.org/contributor-roles/writing-original-draft/"/>
<role content-type="https://credit.niso.org/contributor-roles/writing-review-editing/"/>
<role content-type="https://credit.niso.org/contributor-roles/methodology/"/>
<role content-type="https://credit.niso.org/contributor-roles/software/"/>
<role content-type="https://credit.niso.org/contributor-roles/validation/"/>
<role content-type="https://credit.niso.org/contributor-roles/visualization/"/>
</contrib>
<contrib contrib-type="author">
<name><surname>Jankovsk&#x000FD;</surname> <given-names>Martin</given-names></name>
<xref ref-type="aff" rid="aff1"><sup>1</sup></xref>
<uri xlink:href="http://loop.frontiersin.org/people/2317090/overview"/>
<role content-type="https://credit.niso.org/contributor-roles/writing-original-draft/"/>
<role content-type="https://credit.niso.org/contributor-roles/writing-review-editing/"/>
<role content-type="https://credit.niso.org/contributor-roles/conceptualization/"/>
<role content-type="https://credit.niso.org/contributor-roles/data-curation/"/>
<role content-type="https://credit.niso.org/contributor-roles/formal-analysis/"/>
<role content-type="https://credit.niso.org/contributor-roles/methodology/"/>
</contrib>
<contrib contrib-type="author">
<name><surname>Ju&#x00161;ko</surname> <given-names>Vladim&#x000ED;r</given-names></name>
<xref ref-type="aff" rid="aff2"><sup>2</sup></xref>
<uri xlink:href="http://loop.frontiersin.org/people/2792086/overview"/>
<role content-type="https://credit.niso.org/contributor-roles/writing-original-draft/"/>
<role content-type="https://credit.niso.org/contributor-roles/writing-review-editing/"/>
</contrib>
</contrib-group>
<aff id="aff1"><sup>1</sup><institution>Faculty of Forestry and Wood Sciences, Czech University of Life Sciences Prague</institution>, <addr-line>Prague</addr-line>, <country>Czechia</country></aff>
<aff id="aff2"><sup>2</sup><institution>Faculty of Forestry, Technical University in Zvolen</institution>, <addr-line>Zvolen</addr-line>, <country>Slovakia</country></aff>
<author-notes>
<fn fn-type="edited-by"><p>Edited by: Ken Byrne, University of Limerick, Ireland</p></fn>
<fn fn-type="edited-by"><p>Reviewed by: Tolga Ozturk, Istanbul University-Cerrahpasa, T&#x000FC;rkiye</p>
<p>Francesco Latterini, Polish Academy of Sciences, Poland</p></fn>
<corresp id="c001">&#x0002A;Correspondence: Martin Duchan <email>duchan&#x00040;fld.czu.cz</email></corresp>
<fn fn-type="other" id="fn001"><p>&#x02020;ORCID: Zuzana Dud&#x000E1;kov&#x000E1; <ext-link ext-link-type="uri" xlink:href="https://orcid.org/0000-0003-4776-6903">orcid.org/0000-0003-4776-6903</ext-link></p></fn></author-notes>
<pub-date pub-type="epub">
<day>30</day>
<month>09</month>
<year>2024</year>
</pub-date>
<pub-date pub-type="collection">
<year>2024</year>
</pub-date>
<volume>7</volume>
<elocation-id>1464140</elocation-id>
<history>
<date date-type="received">
<day>13</day>
<month>07</month>
<year>2024</year>
</date>
<date date-type="accepted">
<day>09</day>
<month>09</month>
<year>2024</year>
</date>
</history>
<permissions>
<copyright-statement>Copyright &#x000A9; 2024 Allman, Dud&#x000E1;kov&#x000E1;, Duchan, Jankovsk&#x000FD; and Ju&#x00161;ko.</copyright-statement>
<copyright-year>2024</copyright-year>
<copyright-holder>Allman, Dud&#x000E1;kov&#x000E1;, Duchan, Jankovsk&#x000FD; and Ju&#x00161;ko</copyright-holder>
<license xlink:href="http://creativecommons.org/licenses/by/4.0/"><p>This is an open-access article distributed under the terms of the Creative Commons Attribution License (CC BY). The use, distribution or reproduction in other forums is permitted, provided the original author(s) and the copyright owner(s) are credited and that the original publication in this journal is cited, in accordance with accepted academic practice. No use, distribution or reproduction is permitted which does not comply with these terms.</p></license>
</permissions>
<abstract>
<sec>
<title>Introduction</title>
<p>Timber harvesting on steep terrain is a challenge in terms of economic viability, safety, and environmental performance. Felling with chainsaws and use of yarders seems optimal in this environment. However, using mobile traction winches allows for the safe work of ground-based technologies even in these challenging conditions.</p></sec>
<sec>
<title>Methods</title>
<p>Our study assessed the impact of winch-assisted cut-to-length harvesting on soil disturbance in young forest stands (up to 40 years old) across slopes of 14.9&#x000B0;-27.4&#x000B0; (27&#x02013;52%). Utilizing 78 measurement points (i.e., 234 measurements), we analyzed soil samples from trail ruts, between ruts, and undisturbed areas for soil bulk density (g.cm<sup>&#x02212;3</sup>) and soil moisture content (%), simultaneously measuring penetration resistance (MPa), penetration depth (cm) and rut depth (cm).</p></sec>
<sec>
<title>Results</title>
<p>The results highlighted that areas without winch assistance experienced the most significant increases in soil bulk density (up to 22.35%) and penetration resistance (up to 26.8%), though these differences were not statistically significant. Linear mixed effects models did not confirm a significant effect (<italic>p</italic> &#x0003E; 0.05) of a traction winch on the soil bulk density (g cm<sup>&#x02212;3</sup>) and penetration resistance (MPa) in the ruts of the forwarding trails. Mean forwarding trail profile depths ranged from 4.63 to 7.28 cm, with the maximum depths between 10.86 and 17.25 cm, showing deeper ruts in non-assisted areas. Moreover, the presence of the traction winch (<italic>p</italic> &#x0003C; 0.05) significantly affected the maximal depth of the forwarding trail rut.</p></sec>
<sec>
<title>Conclusion</title>
<p>The findings suggest that winch-assisted harvesting may mitigate soil disturbance (rut depths) on steep slopes, offering a sustainable option for utilizing ground-based machinery with reduced environmental impact.</p></sec></abstract>
<kwd-group>
<kwd>CTL technologies</kwd>
<kwd>mobile traction winch</kwd>
<kwd>steep slopes</kwd>
<kwd>bulk density</kwd>
<kwd>penetration resistance</kwd>
<kwd>soil disturbance</kwd>
</kwd-group>
<counts>
<fig-count count="3"/>
<table-count count="8"/>
<equation-count count="2"/>
<ref-count count="44"/>
<page-count count="12"/>
<word-count count="8896"/>
</counts>
<custom-meta-wrap>
<custom-meta>
<meta-name>section-at-acceptance</meta-name>
<meta-value>Forest Soils</meta-value>
</custom-meta>
</custom-meta-wrap>
</article-meta>
</front>
<body>
<sec id="s1">
<title>1 Introduction</title>
<p>Slovakia is a mountainous country in the Central Europe. Its forest cover reaches 41.3%, which makes it the 13th most forested European country (Ministry of Agriculture Rural Development of the Slovak Republic, <xref ref-type="bibr" rid="B27">2022</xref>). The demanding terrain conditions substantially limit the machinery and technologies Slovak foresters use. Approximately 22% of forests (422,000 ha) are located on slopes over 50% (26.57&#x000B0;), making them impassable for conventional skidders (Ministry of Agriculture Rural Development of the Slovak Republic, <xref ref-type="bibr" rid="B27">2022</xref>). Forest operations in mountainous conditions with steep slopes and rough terrain are particularly challenging (Latterini et al., <xref ref-type="bibr" rid="B23">2024b</xref>). To carry out forest harvesting on steep slopes, foresters should use yarders combined with chainsaw felling (Cavalli, <xref ref-type="bibr" rid="B6">2012</xref>; Holzleitner et al., <xref ref-type="bibr" rid="B15">2018</xref>). Additionally, the forests on steep slopes are frequently young, established after windstorms and bark beetle disturbances (Sommerfeld et al., <xref ref-type="bibr" rid="B37">2021</xref>). Small stem volume and the need to frequently remount the yarders in extreme conditions (Stampfer et al., <xref ref-type="bibr" rid="B38">2006</xref>) contribute to the disinterest of foresters in using cable yarders. Yarders also show lower productivity, higher costs, and higher occupational health risks than ground-based technologies (Holzfeind et al., <xref ref-type="bibr" rid="B14">2018</xref>; Raymond, <xref ref-type="bibr" rid="B32">2010</xref>; Stampfer et al., <xref ref-type="bibr" rid="B38">2006</xref>; Tsioras et al., <xref ref-type="bibr" rid="B40">2014</xref>).</p>
<p>There is a possibility of deployment of cut-to-length (CTL) technologies, which are limited by the slope accessibility of standard forwarders and harvesters (Holzfeind et al., <xref ref-type="bibr" rid="B14">2018</xref>). The highest level of soil disturbance generally occurs when ground-based forest operations are performed on slopes over 20% (Latterini et al., <xref ref-type="bibr" rid="B24">2023</xref>). Cavalli and Amishev (<xref ref-type="bibr" rid="B7">2019</xref>) state that wheeled machinery equipped with wheel chains or semi-tracks can usually access a slope of 45% (24.23&#x000B0;), while tracked machines can climb up to 60% (30.96&#x000B0;). However, if winches secure the machines, they should be able to operate up to a slope between 75% (36.87&#x000B0;) and 85% (40.36&#x000B0;). Visser and Stampfer (<xref ref-type="bibr" rid="B42">2015</xref>) state that for downhill skidding, wheeled skidders can reach up to 50% (26.57&#x000B0;) slopes and crawler tractors can reach up to 60% (30.96&#x000B0;) slopes. Further, the authors state that operating from 50 (26.57&#x000B0;) to 60% (30.96&#x000B0;) slope is a critical zone where purpose-built steep terrain harvesters are required. Indeed, operating above 60% (30.96&#x000B0;) is considered hazardous and requires additional securing systems such as cable-assist or traction winch technology. Similarly, Garren et al. (<xref ref-type="bibr" rid="B10">2019</xref>) state that cable-assist technologies are a viable option for increasing the slope range of ground-based harvesting machinery.</p>
<p>Although there are technological advantages to increasing the slope range of ground-based machinery, the effects of enabling machine traffic in such demanding conditions on forest ecosystems, and especially on forest soils, should not be neglected (Picchio et al., <xref ref-type="bibr" rid="B31">2020</xref>; Jourgholami et al., <xref ref-type="bibr" rid="B19">2014</xref>). The ongoing climate change pressures foresters to assess the anthropic effects on ecosystems and minimize the short- and long-term ecosystem disturbances, such as post-harvesting disturbance. Indeed, through soil and tree damage, such disturbances further reduce the resilience of forest ecosystems to climate change. During forest operations, soil disturbance mainly relates to soil compaction after the machine&#x00027;s passage (Latterini et al., <xref ref-type="bibr" rid="B22">2024a</xref>). CTL machines are heavy, which is a growing problem as producers continuously make larger and heavier machinery (Varol et al., <xref ref-type="bibr" rid="B41">2020</xref>). These heavy forest machines compact the soil when they operate, create ruts (Cambi et al., <xref ref-type="bibr" rid="B4">2015</xref>; Goutal et al., <xref ref-type="bibr" rid="B11">2013</xref>) and lead to topsoil removal and displacement (Horn et al., <xref ref-type="bibr" rid="B16">2004</xref>; Najafi and Solgi, <xref ref-type="bibr" rid="B30">2010</xref>). The compaction of soil reduces its porosity and changes its structure (Macr&#x000ED; et al., <xref ref-type="bibr" rid="B25">2017</xref>; Sohrabi et al., <xref ref-type="bibr" rid="B35">2022</xref>; Solgi et al., <xref ref-type="bibr" rid="B36">2015</xref>). These structural changes then cause an increase in soil penetration resistance, soil bulk density, and soil shear strength (Allman et al., <xref ref-type="bibr" rid="B1">2015</xref>; Cambi et al., <xref ref-type="bibr" rid="B5">2017</xref>; Picchio et al., <xref ref-type="bibr" rid="B31">2020</xref>; Williamson and Neilsen, <xref ref-type="bibr" rid="B44">2000</xref>). Subsequently, it decreases site productivity because soil compaction exceeding three MPa of penetration resistance may hinder root growth (Blouin et al., <xref ref-type="bibr" rid="B3">2008</xref>; Labelle et al., <xref ref-type="bibr" rid="B20">2019</xref>). Wheels or tracks create compacted ruts, where water infiltration capacity is reduced, resulting in increased surface runoff (Ares et al., <xref ref-type="bibr" rid="B2">2005</xref>; Lang et al., <xref ref-type="bibr" rid="B21">2016</xref>). In steep terrains, surface runoff can cause substantial erosion and the removal of fertile topsoil (Etehadi Abari et al., <xref ref-type="bibr" rid="B9">2017</xref>).</p>
<p>While there is a substantial body of research on harvesting-induced soil disturbance in more favorable conditions where skidders and harvesters commonly operate (Allman et al., <xref ref-type="bibr" rid="B1">2015</xref>; Cambi et al., <xref ref-type="bibr" rid="B5">2017</xref>; Macr&#x000ED; et al., <xref ref-type="bibr" rid="B25">2017</xref>; Picchio et al., <xref ref-type="bibr" rid="B31">2020</xref>; Tavankar et al., <xref ref-type="bibr" rid="B39">2017</xref>), studies on cable-assisted or mobile traction (Green et al., <xref ref-type="bibr" rid="B12">2020</xref>) winched CTL technologies working on steep slopes are scarce. There is a lack of literature on soil disturbance (real-life measurements of soil properties) caused by this kind of technology.</p>
<p>To fill this knowledge gap, we asked: what are the effects of mobile traction winch-assisted CTL machines on soil in steep slopes? We hypothesize that the CTL technology causes significantly higher levels of soil disturbance on steep slopes (in terms of soil bulk density, penetration resistance, and deeper rutting).</p></sec>
<sec id="s2">
<title>2 Material and methods</title>
<sec>
<title>2.1 Study area</title>
<p>Forests in the study were managed by Military Forests and Estates of the Slovak Republic, Forest district Ke&#x0017E;marok (<xref ref-type="fig" rid="F1">Figure 1</xref>; GPS coordinates: 49&#x000B0;15&#x02032;12.5&#x02033;N 20&#x000B0;31&#x02032;47.98&#x02033;E). The Forest district was located in the northern part of Slovakia, in the steep, rugged slopes of Levo&#x0010D;a mountain range (Western Carpathians).</p>
<fig id="F1" position="float">
<label>Figure 1</label>
<caption><p>The area of Military Forest Enterprise Ke&#x0017E;marok.</p></caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="ffgc-07-1464140-g0001.tif"/>
</fig>
<p>The Forest district managed 17,833 ha of forests with elevation between 600 and 1,290 m asl. The most abundant species was <italic>Picea abies</italic> L. (63.2%), followed by <italic>Fagus sylvatica</italic> L. (14.4%), <italic>Pinus sylvestris</italic> L. (9.1%), <italic>Abies alba</italic> Mill. (5.8%), <italic>Larix decidua</italic> Mill. (3.5%), with annual harvests reaching &#x0007E;45,000 m<sup>3</sup> of timber and a forest road network density of 8.21 m/ha.</p>
<p>Due to the size and diversity of the area (from the perspective of the natural conditions), we only report on soils in stands with the measurement plots (<xref ref-type="table" rid="T1">Table 1</xref>). Umbric Podsols Schad (<xref ref-type="bibr" rid="B33">2016</xref>) created from lighter solid products of weathering of acidic bedrock were the soil type in observed forest stands. According to soil particle size and texture classes, the soils were sandy loams, with a moderate soil water retention capacity (95&#x02013;130 l/m<sup>2</sup>) and a moderate soil infiltration capacity (0.081&#x02013;0.8 cm/min). The bedrock was created mainly from flyshoid sandstones. Mean annual rainfall in the observed mountain region ranges between 800 and 900 mm, and the mean annual temperature varies between 4 and 6&#x000B0;C. The mean potential evapotranspiration ranges between 400 and 450 mm. Detailed climatic data from the year 2022 were recorded by a GARNI 1025 Arcus (GARNI technology, Ostrava, Czech Republic) weather station located in Lubica village (GPS: 49&#x000B0;07&#x02032;08.9&#x02033;N and 20&#x000B0;29&#x02032;42.9&#x02033;E), the closest (16 km) weather station to the study area. According to the weather station, 595 mm of rainfall (monthly: &#x000D8; 49.55; min. 11.40; max. 107.72) occurred in 2022, which is slightly above normal (12 mm) compared to long-term rainfall (583 mm) at the given weather station. However, during August and September, when the harvesting and field work were conducted, the area experienced a rainfall deficit (August: &#x02212;25.25%; September: &#x0002B;5.02%).</p>
<table-wrap position="float" id="T1">
<label>Table 1</label>
<caption><p>Basic spatial and mensurational information on the observed forest stands.</p></caption>
<table frame="box" rules="all">
<thead>
<tr style="background-color:#919498;color:#ffffff">
<th valign="top" align="center"><bold>Stand</bold></th>
<th valign="top" align="center"><bold>1</bold></th>
<th valign="top" align="center"><bold>2</bold></th>
<th valign="top" align="center"><bold>3</bold></th>
<th valign="top" align="center"><bold>Control</bold></th>
</tr>
</thead>
<tbody>
<tr>
<td valign="top" align="center">GPS coordinates</td>
<td valign="top" align="center">49&#x000B0;13&#x02032;32.1&#x02033;N 20&#x000B0;37&#x02032;45.0&#x02033;E</td>
<td valign="top" align="center">49&#x000B0;13&#x02032;13.5&#x02033;N 20&#x000B0;37&#x02032;43.4&#x02033;E</td>
<td valign="top" align="center">49&#x000B0;13&#x02032;07.7&#x02033;N 20&#x000B0;37&#x02032;56.0&#x02033;E</td>
<td valign="top" align="center">49&#x000B0;13&#x02032;25.7&#x02033;N 20&#x000B0;37&#x02032;27.5&#x02033;E</td>
</tr> <tr>
<td valign="top" align="center">Stand area (ha)</td>
<td valign="top" align="center">15.53</td>
<td valign="top" align="center">7.42</td>
<td valign="top" align="center">17.22</td>
<td valign="top" align="center">7.03</td>
</tr> <tr>
<td valign="top" align="center">Harvested area (ha)</td>
<td valign="top" align="center">9.31</td>
<td valign="top" align="center">4.03</td>
<td valign="top" align="center">8.67</td>
<td valign="top" align="center">2.89</td>
</tr> <tr>
<td valign="top" align="center">Harvested volume by tree species (m<sup>3</sup>)</td>
<td valign="top" align="center">Pa:442.17 Fs: 1.28</td>
<td valign="top" align="center">Pa:167.38 Fs: 54.41</td>
<td valign="top" align="center">Pa:391.29 Aa: 13.13</td>
<td valign="top" align="center">Pa: 109.34</td>
</tr> <tr>
<td valign="top" align="center">Harvested volume &#x02211; (m<sup>3</sup>)</td>
<td valign="top" align="center">443.45</td>
<td valign="top" align="center">221.79</td>
<td valign="top" align="center">404.42</td>
<td valign="top" align="center">109.34</td>
</tr> <tr>
<td valign="top" align="center">Harvested volume (m<sup>3</sup>/ha)</td>
<td valign="top" align="center">47.63</td>
<td valign="top" align="center">55.04</td>
<td valign="top" align="center">46.65</td>
<td valign="top" align="center">37.83</td>
</tr> <tr>
<td valign="top" align="center">Stand age (years)</td>
<td valign="top" align="center">30</td>
<td valign="top" align="center">35</td>
<td valign="top" align="center">40</td>
<td valign="top" align="center">30</td>
</tr> <tr>
<td valign="top" align="center">Tree species composition (%)</td>
<td valign="top" align="center">Pa: 85 Fs: 15</td>
<td valign="top" align="center">Pa: 70 Fs: 30</td>
<td valign="top" align="center">Pa: 85 Fs: 10 Aa: 5</td>
<td valign="top" align="center">Pa: 95 Aa: 5</td>
</tr> <tr>
<td valign="top" align="center">Mean volume of harvested stems (m<sup>3</sup>)<sup>a)</sup></td>
<td valign="top" align="center">Pa: 0.11 Fs: 0.12</td>
<td valign="top" align="center">Pa: 0.28 Fs: 0.23</td>
<td valign="top" align="center">Pa: 0.20 Fs: 0.27 Aa: 0.21</td>
<td valign="top" align="center">Pa: 0.25 Aa: 0.35</td>
</tr> <tr>
<td valign="top" align="center">Stocking</td>
<td valign="top" align="center">0.85</td>
<td valign="top" align="center">0.85</td>
<td valign="top" align="center">0.85</td>
<td valign="top" align="center">0.75</td>
</tr> <tr>
<td valign="top" align="center">Elevation (m asl)</td>
<td valign="top" align="center">970&#x02013;1,020</td>
<td valign="top" align="center">940&#x02013;1,050</td>
<td valign="top" align="center">870&#x02013;1,010</td>
<td valign="top" align="center">890&#x02013;1,050</td>
</tr> <tr>
<td valign="top" align="center">Mean slope % (&#x000B0;)</td>
<td valign="top" align="center">51.8 (27.4&#x000B0;)</td>
<td valign="top" align="center">46.4 (24.9&#x000B0;)</td>
<td valign="top" align="center">50.5 (26.8&#x000B0;)</td>
<td valign="top" align="center">26.7 (14.9&#x000B0;)</td>
</tr> <tr>
<td valign="top" align="center">Exposure</td>
<td valign="top" align="center">East</td>
<td valign="top" align="center">East</td>
<td valign="top" align="center">East</td>
<td valign="top" align="center">North-west</td>
</tr> <tr>
<td valign="top" align="center">Forwarding distance (m)</td>
<td valign="top" align="center">700</td>
<td valign="top" align="center">700</td>
<td valign="top" align="center">300</td>
<td valign="top" align="center">1,000</td>
</tr> <tr>
<td valign="top" align="center">Number of sample plots/measurement locations</td>
<td valign="top" align="center">12/24</td>
<td valign="top" align="center">10/20</td>
<td valign="top" align="center">11/22</td>
<td valign="top" align="center">6/12</td>
</tr> <tr>
<td valign="top" align="center">Technology with traction assistance</td>
<td valign="top" align="center">Yes</td>
<td valign="top" align="center">Yes</td>
<td valign="top" align="center">Yes</td>
<td valign="top" align="center">No</td>
</tr></tbody>
</table>
<table-wrap-foot>
<p><sup><italic>a</italic>)</sup>Pa, <italic>Picea abies</italic> (L.); Fs, Fagus <italic>sylvatica</italic> (L.); Aa, <italic>Abies alba</italic> (Mill).</p>
</table-wrap-foot>
</table-wrap>
</sec>
<sec>
<title>2.2 Logging operations and technology</title>
<p>The machinery was used as follows: a ROTTNE H11<sup>C</sup> harvester and a ROTTNE F15<sup>D</sup> forwarder were used to fell, limb, buck, and extract timber from the forest stand to the roadside while secured by two Ecoforst TWINCH 10.1 mobile traction winches (i.e., a single winch per machine), located on the hilltops and radio-controlled by the harvester and forwarder operators (<xref ref-type="table" rid="T2">Table 2</xref>). The harvester carried out thinning operations and passed over 4 m wide trails, which the operator created (<xref ref-type="supplementary-material" rid="SM1">Supplementary Figure 1</xref>). Winch-assisted harvesting took place in stands with steep slopes (46.4&#x02013;51.8%) no. 1, 2, and 3. The CTL machinery worked without a winch-assist in the Control stand because the slope was not steep (26.7%) and enabled safe traffic (<xref ref-type="table" rid="T1">Table 1</xref>).</p>
<table-wrap position="float" id="T2">
<label>Table 2</label>
<caption><p>Basic technical parameters of the machines deployed in the observed forest stands.</p></caption>
<table frame="box" rules="all">
<thead>
<tr style="background-color:#919498;color:#ffffff">
<th valign="top" align="left"><bold>Machine</bold></th>
<th valign="top" align="center"><bold>Rottne H11<sup>C</sup></bold></th>
<th valign="top" align="center"><bold>Rottne F15<sup>D</sup></bold></th>
<th valign="top" align="center"><bold>Machine</bold></th>
<th valign="top" align="center"><bold>Ecoforst TWINCH 10.1</bold></th>
</tr>
</thead>
<tbody>
<tr>
<td valign="top" align="left">Type of machine</td>
<td valign="top" align="center">Harvester</td>
<td valign="top" align="center">Forwarder</td>
<td valign="top" align="center">Type of machine</td>
<td valign="top" align="center">Mobile traction winch</td>
</tr> <tr>
<td valign="top" align="left">Engine output at 1,800 rpm (kW)</td>
<td valign="top" align="center">164</td>
<td valign="top" align="center">164</td>
<td valign="top" align="center">Diesel engine/power (kW)</td>
<td valign="top" align="center">IVECO S30 ENT&#x02014;V/107</td>
</tr> <tr>
<td valign="top" align="left">Tractive force (kN)</td>
<td valign="top" align="center">167</td>
<td valign="top" align="center">177</td>
<td valign="top" align="center">Fuel tank (l)</td>
<td valign="top" align="center">280</td>
</tr> <tr>
<td valign="top" align="left">Crane</td>
<td valign="top" align="center">RK 185</td>
<td valign="top" align="center">RK 125</td>
<td valign="top" align="center">Weight (kg)</td>
<td valign="top" align="center">7,800</td>
</tr> <tr>
<td valign="top" align="left">Lifting torque/torque (kNm)</td>
<td valign="top" align="center">193/50.8</td>
<td valign="top" align="center">125/35.8</td>
<td valign="top" align="center">Pressure on the soil surface (kg/cm<sup>2</sup>)</td>
<td valign="top" align="center">0.45</td>
</tr> <tr>
<td valign="top" align="left">Service weight (kg)</td>
<td valign="top" align="center">18,900</td>
<td valign="top" align="center">18,700</td>
<td valign="top" align="center">Maximum pulling force (kN)</td>
<td valign="top" align="center">80</td>
</tr> <tr>
<td valign="top" align="left">Wheels</td>
<td valign="top" align="center">8WD</td>
<td valign="top" align="center">8WD</td>
<td valign="top" align="center">Rope diameter (mm)</td>
<td valign="top" align="center">18.5</td>
</tr> <tr>
<td valign="top" align="left">Tire</td>
<td valign="top" align="center">Nokian 710/40&#x02013;24.5</td>
<td valign="top" align="center">Nokian 710/45&#x02013;26.5</td>
<td valign="top" align="center">Rope length (m)</td>
<td valign="top" align="center">500</td>
</tr> <tr>
<td valign="top" align="left">Tire inflation pressure (kPa)</td>
<td valign="top" align="center">500</td>
<td valign="top" align="center">550</td>
<td/>
<td/>
</tr> <tr>
<td valign="top" align="left">Tracks</td>
<td valign="top" align="center">Clark tracks HX&#x02212;24&#x02013;75</td>
<td valign="top" align="center">Clark tracks HX&#x02212;26&#x02013;75</td>
<td valign="top" align="center">Maximum speed (km/h)</td>
<td valign="top" align="center">4.0</td>
</tr> <tr>
<td valign="top" align="left">Operators age (years)</td>
<td valign="top" align="center">42</td>
<td valign="top" align="center">38</td>
<td valign="top" align="left" colspan="2">Operators are the same</td>
</tr> <tr>
<td valign="top" align="left">Operators experience (years)</td>
<td valign="top" align="center">7</td>
<td valign="top" align="center">18</td>
<td/>
<td/>
</tr></tbody>
</table>
</table-wrap>
</sec>
<sec>
<title>2.3 Measurement process and devices</title>
<p>We conducted the measurements between October 10 and 15, 2022, immediately after the harvesting machinery left the forest stands, and it was safe to begin measuring. During fieldwork, no precipitation occurred, and temperature ranged between 7 and 15&#x000B0;C. Forest stands were selected based on their similarity in soils, slopes, tree species mix, tree spatial distribution, other natural conditions, and proximity (<xref ref-type="table" rid="T1">Table 1</xref>; <xref ref-type="fig" rid="F2">Figure 2</xref>). Sample plots were established on particular forwarding trails. The sample plot size was 10% of the harvested area in stands up to 50,000 m<sup>2</sup> or 5% in stands where the harvested area was larger than 50,000 m<sup>2</sup> (Allman et al., <xref ref-type="bibr" rid="B1">2015</xref>). Plots were squares, with a side of 20 &#x000D7; 20 m. The positioning of the sample plots was calculated according to Scheer and Sedm&#x000E1;k (<xref ref-type="bibr" rid="B34">2010</xref>) (<xref ref-type="disp-formula" rid="E1">Equation 1</xref>):</p>
<disp-formula id="E1"><label>(1)</label><mml:math id="M1"><mml:mtable class="eqnarray" columnalign="left"><mml:mtr><mml:mtd><mml:mi>s</mml:mi><mml:mo>=</mml:mo><mml:mn>100</mml:mn><mml:mo>&#x000D7;</mml:mo><mml:msqrt><mml:mrow><mml:mfrac><mml:mrow><mml:mi>P</mml:mi></mml:mrow><mml:mrow><mml:mi>n</mml:mi></mml:mrow></mml:mfrac></mml:mrow></mml:msqrt></mml:mtd></mml:mtr></mml:mtable></mml:math></disp-formula>
<p>s, sample plot distance (m)</p>
<p><italic>P</italic>, harvested area (ha)</p>
<p><italic>n</italic>, number of sample plots</p>
<p>Measurement locations were established on the opposite sides of each plot (<xref ref-type="fig" rid="F3">Figure 3</xref>). In each measurement location, we (i) determined the longitudinal slope with a True Pulse 360B (Laser Tech Inc., Colorado, USA) laser range finder (in &#x000B0;); (ii) evaluated the presence and depth of slash on the trail (cm), (iii) measured the forwarding trail profile (cm) and maximal rut depth, (iv) collected soil samples from the localities stand, rut and in the middle of the trail to determine the bulk density (BD) of soil (g cm<sup>&#x02212;3</sup>) and soil moisture content (%), (v) measured soil penetration resistance (PR) in MPa and penetration depth (cm).</p>
<fig id="F2" position="float">
<label>Figure 2</label>
<caption><p>Distance and position of the forest stands where measurements took place.</p></caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="ffgc-07-1464140-g0002.tif"/>
</fig>
<fig id="F3" position="float">
<label>Figure 3</label>
<caption><p>Measurement locations on the opposite sides of sample plots on skid trails.</p></caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="ffgc-07-1464140-g0003.tif"/>
</fig>
<p>To measure the average forwarding trail profile depth (cm), we placed a 4 m ranging rod perpendicularly on the trail. We measured the trail profile with a tape measure from the soil surface to the lower side of the rod, each 20 cm of the length of the ranging rod (Jankovsk&#x000FD; et al., <xref ref-type="bibr" rid="B17">2019</xref>). Maximal rut depth (cm) was expressed as the largest value measured in the ruts of the forwarding trail. The intensity of disturbance via rut creation was classified according to the rut disturbance classification scale, shown in <xref ref-type="table" rid="T3">Table 3</xref>. Soil samples were collected into 100 cm<sup>3</sup> Eijkelkamp sampling cylinders (Giesbeek, Netherlands). Immediately after sampling, the cylinders were sealed in a plastic bag for subsequent BD and moisture content laboratory analyses. BD of soil and moisture content (<xref ref-type="disp-formula" rid="E2">Equation 2</xref>) was determined in a dried state from samples dried for 24 h at 105&#x000B0;C (Allman et al., <xref ref-type="bibr" rid="B1">2015</xref>).</p>
<disp-formula id="E2"><label>(2)</label><mml:math id="M2"><mml:mtable class="eqnarray" columnalign="left"><mml:mtr><mml:mtd><mml:mi>M</mml:mi><mml:mi>o</mml:mi><mml:mi>i</mml:mi><mml:mi>s</mml:mi><mml:mi>t</mml:mi><mml:mi>u</mml:mi><mml:mi>r</mml:mi><mml:mi>e</mml:mi><mml:mtext>&#x000A0;</mml:mtext><mml:mi>c</mml:mi><mml:mi>o</mml:mi><mml:mi>n</mml:mi><mml:mi>t</mml:mi><mml:mi>e</mml:mi><mml:mi>n</mml:mi><mml:mi>t</mml:mi><mml:mtext>&#x000A0;</mml:mtext><mml:mrow><mml:mo stretchy="false">(</mml:mo><mml:mrow><mml:mi>%</mml:mi></mml:mrow><mml:mo stretchy="false">)</mml:mo></mml:mrow><mml:mo>=</mml:mo><mml:mtext>&#x000A0;</mml:mtext><mml:mrow><mml:mo stretchy="false">(</mml:mo><mml:mrow><mml:mfrac><mml:mrow><mml:mi>r</mml:mi><mml:mi>a</mml:mi><mml:mi>w</mml:mi><mml:mtext>&#x000A0;</mml:mtext><mml:mi>s</mml:mi><mml:mi>a</mml:mi><mml:mi>m</mml:mi><mml:mi>p</mml:mi><mml:mi>l</mml:mi><mml:mi>e</mml:mi><mml:mtext>&#x000A0;</mml:mtext><mml:mrow><mml:mo stretchy="false">(</mml:mo><mml:mrow><mml:mi>g</mml:mi></mml:mrow><mml:mo stretchy="false">)</mml:mo></mml:mrow><mml:mo>-</mml:mo><mml:mi>d</mml:mi><mml:mi>r</mml:mi><mml:mi>i</mml:mi><mml:mi>e</mml:mi><mml:mi>d</mml:mi><mml:mtext>&#x000A0;</mml:mtext><mml:mi>s</mml:mi><mml:mi>a</mml:mi><mml:mi>m</mml:mi><mml:mi>p</mml:mi><mml:mi>l</mml:mi><mml:mi>e</mml:mi><mml:mtext>&#x000A0;</mml:mtext><mml:mrow><mml:mo stretchy="false">(</mml:mo><mml:mrow><mml:mi>g</mml:mi></mml:mrow><mml:mo stretchy="false">)</mml:mo></mml:mrow></mml:mrow><mml:mrow><mml:mi>d</mml:mi><mml:mi>r</mml:mi><mml:mi>i</mml:mi><mml:mi>e</mml:mi><mml:mi>d</mml:mi><mml:mtext>&#x000A0;</mml:mtext><mml:mi>s</mml:mi><mml:mi>a</mml:mi><mml:mi>m</mml:mi><mml:mi>p</mml:mi><mml:mi>l</mml:mi><mml:mi>e</mml:mi><mml:mtext>&#x000A0;</mml:mtext><mml:mrow><mml:mo stretchy="false">(</mml:mo><mml:mrow><mml:mi>g</mml:mi></mml:mrow><mml:mo stretchy="false">)</mml:mo></mml:mrow></mml:mrow></mml:mfrac></mml:mrow><mml:mo stretchy="false">)</mml:mo></mml:mrow><mml:mo>&#x000D7;</mml:mo><mml:mn>100</mml:mn></mml:mtd></mml:mtr></mml:mtable></mml:math></disp-formula>
<p>Penetration resistance and penetration depth of soil were measured in the exact locations of stand, rut and middle by an Eijkelkamp Penetrologger (Giesbeek, Netherlands), equipped with an 80 cm extension rod and a 30&#x000B0; cone screwed onto the bottom end of the rod. The cone had a nominal diameter of 11.28 mm and a base area of 1 cm<sup>2</sup>. Thirty-nine sample plots containing 78 measurement locations were established in total. From the plots, 234 BD and moisture content samples were collected, and 234 PR measurements were carried out. Penetration resistance and penetration depth were measured by an Eijkelkamp Penetrologger, which recorded values per 1 cm of penetration in the range between 0 and 80 cm, based on the depth to which we were able to push the penetrometer which resulted in a higher number of values for these variables.</p>
<table-wrap position="float" id="T3">
<label>Table 3</label>
<caption><p>Rut disturbance classification scale used to observe soil disturbance levels.</p></caption>
<table frame="box" rules="all">
<thead>
<tr style="background-color:#919498;color:#ffffff">
<th valign="top" align="left"><bold>Level of soil disturbance</bold></th>
<th valign="top" align="center"><bold>Rut depth (cm)</bold></th>
<th valign="top" align="left"><bold>Disturbance characteristic</bold></th>
</tr>
</thead>
<tbody>
<tr>
<td valign="top" align="left">No disturbance</td>
<td valign="top" align="center">0</td>
<td valign="top" align="left">Soil with unimpaired vegetation and top humus layer.</td>
</tr> <tr>
<td valign="top" align="left">Very low disturbance</td>
<td valign="top" align="center">0</td>
<td valign="top" align="left">Humus layer partially removed (&#x0003C; 50%) or mixed with humus soil horizon. Soil is without any visible signs of compression. Soil cover damaged.</td>
</tr> <tr>
<td valign="top" align="left">Low disturbance</td>
<td valign="top" align="center">&#x0003C; 7.9</td>
<td valign="top" align="left">The humus layer removed. Humus horizon partially (&#x0003C; 50%) removed. Soil compressed by machinery passage in low precipitation conditions. Bearing capacity unimpaired.</td>
</tr> <tr>
<td valign="top" align="left">Moderate disturbance</td>
<td valign="top" align="center">8&#x02013;15.9</td>
<td valign="top" align="left">Top soil horizon removed partially (50&#x02013;80%). Soil compressed, water permeability decreased signifi cantly. Visible lateral soil displacement near ruts.</td>
</tr> <tr>
<td valign="top" align="left">Severe disturbance</td>
<td valign="top" align="center">16&#x02013;25.9</td>
<td valign="top" align="left">Mineral soil partially removed. Soil compressed or destructed by the tires, laterally displaced. Low to very low water permeability of soil. Water stays on the surface of ruts.</td>
</tr> <tr>
<td valign="top" align="left">Very severe disturbance</td>
<td valign="top" align="center">&#x02265;26</td>
<td valign="top" align="left">Mineral horizon uncovered. Very deep ruts, soil in ruts impermeable to water</td>
</tr></tbody>
</table>
</table-wrap>
</sec>
<sec>
<title>2.4 Statistical analyses</title>
<p>Data were processed in Tibco Statistica 14.0.1 and IBM SPSS statistical software. Shapiro-Wilk and Levene&#x00027;s tests were used to test the normality of data distribution and the homogeneity of variance for BD, moisture content, PR, and penetration depth.</p>
<p>One-way analysis of variance (ANOVA) was used to compare the differences between the forest stands (Duncan&#x00027;s test) and measurement locations (Tukey&#x00027;s test) for BD, soil moisture, PR and penetration depth, as well as to compare the differences between the mean and maximal rut depths formed in the observed forest stands. Linear mixed effects models (LMMs) were used to identify the effects of a particular forest stand (random effect) and measurement location (fixed effect) on the variability of BD, soil moisture, PR, and penetration depth, as well as to identify the effects of the presence of traction winches, slash in the forwarding trails, slope, and soil moisture content as fixed effects on BD, PR, and maximal rut depth in forwarding trails as dependent variables. Furthermore, we used LMMs to observe the relationships between PR and BD and the thickness of the slash cover and PR and BD on data from the rut location. We hypothesize that CTL machines cause significantly higher H<sub>1</sub> soil compaction in the ruts, expressed as BD; H<sub>2</sub> penetration resistances (PR) in the ruts of the forwarding trails; H<sub>3</sub> rut depths in forwarding trails (cm), compared to CTL machines deployed in standard conditions without winch assistance. The hypotheses H<sub>1</sub>, H<sub>2</sub>, and H<sub>3</sub> were tested on a 5% &#x003B1; level of significance, i.e., they were rejected, and the alternative hypothesis H<sub>A</sub> &#x0201C;no significant differences between the variables were confirmed&#x0201D; was accepted.</p></sec>
</sec>
<sec id="s3">
<title>3 Results</title>
<p>Shapiro&#x02013;Wilk&#x00027;s test showed that data were not normally distributed at the particular measurement locations.</p>
<sec>
<title>3.1 Bulk density and soil moisture</title>
<p>From the mean dried BD values, it can be seen that machine traffic increased BD in the rut by 13.42 (no. 2)&#x02212;22.35% (Control), representing the most substantial BD increase from all observed forest stands. In contrast, BD was lower in the middle (0.77&#x02013;0.86 g.cm<sup>&#x02212;3</sup>) than in the stand (0.82&#x02013;0.90 g.cm<sup>&#x02212;3</sup>), probably due to the soil displacement into the trail&#x00027;s center caused by machine traffic. The lowest BD values in the ruts were recorded in stands no. 2 (0.93 g.cm<sup>&#x02212;3</sup>) and Control (1.04 g.cm<sup>&#x02212;3</sup>), where the lowest amount of timber was harvested (221.79 and 109.34 m<sup>3</sup>, respectively), resulting in smaller passages of forest operations over the trails. The highest values of moisture content (%) were recorded at the rut location in all forest stands, except for stand no. 3, and fluctuated in the range of 44.24 (no. 1)&#x02212;54.29 (no. 2) % (<xref ref-type="table" rid="T4">Table 4</xref>). The machine passage and the resulting change in the microrelief in the trails caused higher moisture content (%) in the ruts. One-way ANOVA (Duncan&#x00027;s test) did not confirm significant differences in BD between the forest stands (no. 1, 2, 3, Control) at the exact measurement locations, i.e., stand (<italic>F</italic> = 0.638; <italic>p</italic> = 0.593), rut (<italic>F</italic> = 2.373; <italic>p</italic> = 0.077), and middle (<italic>F</italic> = 1.553; <italic>p</italic> = 0.208). The results showed that the conditions and the intensity of soil disturbance in the ruts and middle locations were similar between the forest stands. The differences between locations (Tukey&#x00027;s test) were not significant only in the case of the Control stand (<italic>F</italic> = 2.587; <italic>p</italic> = 0.09) (<xref ref-type="table" rid="T4">Table 4</xref>). Regarding moisture content, the differences between the four observed forest stands were insignificant only at the rut location (<italic>F</italic> = 2.503; <italic>p</italic> = 0.07). Moreover, in the case of stand no. 3, the differences between the stand, rut, and middle measurement locations were insignificant (<italic>F</italic> = 14.816; <italic>p</italic> = 0.00) (<xref ref-type="table" rid="T4">Table 4</xref>).</p>
<table-wrap position="float" id="T4">
<label>Table 4</label>
<caption><p>Mean bulk density and soil moisture values and their differences (%) analyzed through Tukey&#x00027;s test between measurement locations and observed stand no. (Duncan&#x00027;s test).</p></caption>
<table frame="box" rules="all">
<thead>
<tr style="background-color:#919498;color:#ffffff">
<th/>
<th valign="top" align="center" colspan="2"><bold>Stand (S)</bold></th>
<th valign="top" align="center" colspan="2"><bold>Rut (R)</bold></th>
<th valign="top" align="center" colspan="2"><bold>Middle (M)</bold></th>
<th/>
</tr>
</thead>
<tbody>
<tr style="background-color:#919498;color:#ffffff">
<td valign="top" align="center"><bold>Stand no</bold>.</td>
<td valign="top" align="center"><bold>&#x000D8;</bold> BD<sup>a)</sup> &#x000B1;<italic>SD</italic><sup>b)</sup></td>
<td valign="top" align="center">&#x00394;<sup><italic>c</italic>)</sup> <bold>S</bold>&#x02192;<bold>R (%)</bold></td>
<td valign="top" align="center"><bold>&#x000D8; BD</bold> &#x000B1;<bold>SD</bold></td>
<td valign="top" align="center">&#x00394; <bold>R</bold>&#x02192;<bold>M (%)</bold></td>
<td valign="top" align="center"><bold>&#x000D8; BD</bold> &#x000B1;<bold>SD</bold></td>
<td valign="top" align="center">&#x00394; <bold>S</bold>&#x02192;<bold>M (%)</bold></td>
<td valign="top" align="center"><bold>Tukey</bold><sup><italic>e</italic>)</sup> &#x021C6;</td>
</tr> <tr>
<td valign="top" align="center">1</td>
<td valign="top" align="center">0.86 &#x000B1; 0.10</td>
<td valign="top" align="center"><bold>&#x0002B;22.09</bold></td>
<td valign="top" align="center">1.05 &#x000B1; 0.20</td>
<td valign="top" align="center"><bold>&#x02212;18.1</bold></td>
<td valign="top" align="center">0.86 &#x000B1; 0.15</td>
<td valign="top" align="center">0</td>
<td valign="top" align="center">S&#x02013;R; R&#x02013;M</td>
</tr> <tr>
<td valign="top" align="center">2</td>
<td valign="top" align="center">0.82 &#x000B1; 0.12</td>
<td valign="top" align="center">&#x0002B;13.42</td>
<td valign="top" align="center">0.93 &#x000B1; 0.22</td>
<td valign="top" align="center"><bold>&#x02212;17.20</bold></td>
<td valign="top" align="center">0.77 &#x000B1; 0.14</td>
<td valign="top" align="center">&#x02212;6.10</td>
<td valign="top" align="center">R&#x02013;M</td>
</tr> <tr>
<td valign="top" align="center">3</td>
<td valign="top" align="center">0.90 &#x000B1; 0.12</td>
<td valign="top" align="center"><bold>&#x0002B;16.67</bold></td>
<td valign="top" align="center">1.05 &#x000B1; 0.17</td>
<td valign="top" align="center"><bold>&#x02212;19.05</bold></td>
<td valign="top" align="center">0.85 &#x000B1; 0.13</td>
<td valign="top" align="center">&#x02212;5.56</td>
<td valign="top" align="center">S&#x02013;R; R&#x02013;M</td>
</tr> <tr>
<td valign="top" align="center">Control</td>
<td valign="top" align="center">0.85 &#x000B1; 0.38</td>
<td valign="top" align="center">&#x0002B;22.35</td>
<td valign="top" align="center">1.04 &#x000B1; 0.14</td>
<td valign="top" align="center">&#x02212;20.19</td>
<td valign="top" align="center">0.83 &#x000B1; 0.11</td>
<td valign="top" align="center">&#x02212;2.35</td>
<td valign="top" align="center"><italic>p</italic> &#x0003E; 0.05</td>
</tr> <tr>
<td valign="top" align="center">Duncan<sup><italic>f</italic>)</sup> &#x02195;</td>
<td valign="top" align="center"><italic>p</italic> &#x0003E; 0.05</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center"><italic>p</italic> &#x0003E; 0.05</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center"><italic>p</italic> &#x0003E; 0.05</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">&#x02013;</td>
</tr>
<tr style="background-color:#919498;color:#ffffff">
<td valign="top" align="center"><bold>Stand no</bold>.</td>
<td valign="top" align="center"><bold>&#x000D8; MC</bold> &#x000B1;<bold>SD</bold></td>
<td valign="top" align="center">&#x00394; <bold>S</bold>&#x02192;<bold>R (%)</bold></td>
<td valign="top" align="center"><bold>&#x000D8; MC</bold> &#x000B1;<bold>SD</bold></td>
<td valign="top" align="center">&#x00394; <bold>R</bold>&#x02192;<bold>M (%)</bold></td>
<td valign="top" align="center"><bold>&#x000D8;</bold> <italic>MC</italic><sup>d)</sup> &#x000B1;<bold>SD</bold></td>
<td valign="top" align="center">&#x00394; <bold>S</bold>&#x02192;<bold>M (%)</bold></td>
<td valign="top" align="center"><bold>Tukey</bold><sup><italic>e</italic>)</sup> &#x021C6;</td>
</tr> <tr>
<td valign="top" align="center">1</td>
<td valign="top" align="center">34.85 &#x000B1; 11.14</td>
<td valign="top" align="center">&#x0002B;29.94</td>
<td valign="top" align="center">44.24 &#x000B1; 14.90</td>
<td valign="top" align="center">&#x02212;8.97</td>
<td valign="top" align="center">40.27 &#x000B1; 13.10</td>
<td valign="top" align="center">&#x0002B;15.55</td>
<td valign="top" align="center"><italic>p</italic> &#x0003E; 0.05</td>
</tr> <tr>
<td valign="top" align="center">2</td>
<td valign="top" align="center">44.52 &#x000B1; 9.73</td>
<td valign="top" align="center">&#x0002B;21.95</td>
<td valign="top" align="center">54.29 &#x000B1; 25.39</td>
<td valign="top" align="center">&#x02212;4.57</td>
<td valign="top" align="center">51.81 &#x000B1; 12.86</td>
<td valign="top" align="center">&#x0002B;16.38</td>
<td valign="top" align="center"><italic>p</italic> &#x0003E; 0.05</td>
</tr> <tr>
<td valign="top" align="center">3</td>
<td valign="top" align="center">28.82 &#x000B1; 8.10</td>
<td valign="top" align="center"><bold>&#x0002B;40.91</bold></td>
<td valign="top" align="center">40.61 &#x000B1; 8.24</td>
<td valign="top" align="center">&#x0002B;7.46</td>
<td valign="top" align="center">43.64 &#x000B1; 11.82</td>
<td valign="top" align="center"><bold>&#x0002B;51.42</bold></td>
<td valign="top" align="center">S&#x02013;R; S&#x02013;M</td>
</tr> <tr>
<td valign="top" align="center">Control</td>
<td valign="top" align="center">46.51 &#x000B1; 19.50</td>
<td valign="top" align="center">&#x0002B;3.99</td>
<td valign="top" align="center">48.37 &#x000B1; 14.50</td>
<td valign="top" align="center">&#x02212;4.32</td>
<td valign="top" align="center">46.28 &#x000B1; 7.95</td>
<td valign="top" align="center">&#x02212;0.50</td>
<td valign="top" align="center"><italic>p</italic> &#x0003E; 0.05</td>
</tr> <tr>
<td valign="top" align="center">Duncan<sup><italic>f</italic>)</sup> &#x02195;</td>
<td valign="top" align="center">1&#x02013;2; 1&#x02013;Con.; 2&#x02013;3; 3&#x02013;Con.</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center"><italic>p</italic> &#x0003E; 0.05</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">1&#x02013;2</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">&#x02013;</td>
</tr></tbody>
</table>
<table-wrap-foot>
<p><sup><italic>a</italic>)</sup>BD, bulk density (g.cm<sup>&#x02212;3</sup>); <sup><italic>b</italic>)</sup>SD, standard deviation; <sup><italic>c</italic>)</sup>&#x00394;, value difference (%); bold typeface indicates the significance of differences (<italic>p</italic> &#x02264; 0.05) between locations; <sup><italic>d</italic>)</sup>MC, moisture content (%); <sup><italic>e</italic>)</sup>Tukey, between-group comparisons of differences between the measurement locations within each particular forest stand; <sup><italic>f</italic>)</sup>Duncan, between-group comparisons of differences of the same measurement location between particular forest stands.</p>
</table-wrap-foot>
</table-wrap>
<p>Linear mixed model of the effects of measurement locations (stand, rut, middle) as fixed effects, and the forest stand no. (random effect) on BD showed that 18.9% (<inline-formula><mml:math id="M4"><mml:msubsup><mml:mrow><mml:mi>R</mml:mi></mml:mrow><mml:mrow><mml:mtext>m</mml:mtext></mml:mrow><mml:mrow><mml:mn>2</mml:mn></mml:mrow></mml:msubsup></mml:math></inline-formula> = 0.189) of the BD variability was affected by measurement location (marginal effect) and 22.4% (<italic>R</italic><sup>2</sup>c = 0.224) by fixed and random effects (conditional effect). Only about 3.5% of the variability could be attributed to random effects. Measurement location as the fixed effects significantly affected BD (<italic>F</italic> = 28.297; <italic>p</italic> &#x0003C; 0.05). In the case of moisture content as a dependent variable, the <italic>R</italic><sup>2</sup>m was 0.067, and <italic>R</italic><sup>2</sup>c was 0.202, and the random effects (stand no.) explained more variability of the moisture content (13.5%) than fixed effects (6.7%) and had (fixed effect) a significant relationship with moisture content (<italic>F</italic> = 9.749; <italic>p</italic> &#x0003C; 0.05).</p>
<p>We then observed the relationship between the BD in ruts and the usage of traction winches, slope category, and moisture content category as fixed effects and the stand no. as random effects via an LMM (<xref ref-type="table" rid="T5">Table 5</xref>). The analysis showed that 73.2% (<inline-formula><mml:math id="M5"><mml:msubsup><mml:mrow><mml:mi>R</mml:mi></mml:mrow><mml:mrow><mml:mtext>m</mml:mtext></mml:mrow><mml:mrow><mml:mn>2</mml:mn></mml:mrow></mml:msubsup></mml:math></inline-formula> = 0.732) of rut BD variability was explained by fixed effects and 75.7% (<italic>R</italic><sup>2</sup>c = 0.757) by the combination of fixed and random effects (<xref ref-type="table" rid="T5">Table 5</xref>). The random effect (stand no.) contributed only 2.5% to explaining rut BD variability, although only moisture content significantly affected the dependent variable (<italic>F</italic> = 49.164; <italic>p</italic> &#x0003C; 0.05).</p>
<table-wrap position="float" id="T5">
<label>Table 5</label>
<caption><p>Results of testing the fixed effects (traction winch usage, slash cover presence, slope category, moisture content category) for rut bulk density, rut penetration resistance or maximal rut depth, respectively, as dependent variables.</p></caption>
<table frame="box" rules="all">
<thead>
<tr style="background-color:#919498;color:#ffffff">
<th valign="top" align="center"><bold>Source</bold></th>
<th valign="top" align="center"><bold>Numerator df</bold></th>
<th valign="top" align="center"><bold>Denominator df</bold></th>
<th valign="top" align="center"><bold><italic>F</italic></bold></th>
<th valign="top" align="center"><bold>Sig</bold>.</th>
</tr>
</thead>
<tbody>
<tr style="background-color:#dee1e1">
<td valign="top" align="left" colspan="5"><bold>Tests of fixed effects (bulk density g cm</bold><sup>&#x02212;3)</sup></td>
</tr> <tr>
<td valign="top" align="center">Intercept</td>
<td valign="top" align="center">1</td>
<td valign="top" align="center">8.212</td>
<td valign="top" align="center">214.116</td>
<td valign="top" align="center">&#x0003C; 0.001</td>
</tr> <tr>
<td valign="top" align="center">Traction winch</td>
<td valign="top" align="center">1</td>
<td valign="top" align="center">4.323</td>
<td valign="top" align="center">0.280</td>
<td valign="top" align="center">0.623</td>
</tr> <tr>
<td valign="top" align="center">Slash cover</td>
<td valign="top" align="center">1</td>
<td valign="top" align="center">67.102</td>
<td valign="top" align="center">2.056</td>
<td valign="top" align="center">0.156</td>
</tr> <tr>
<td valign="top" align="center">Slope category (%)</td>
<td valign="top" align="center">1</td>
<td valign="top" align="center">6.461</td>
<td valign="top" align="center">0.018</td>
<td valign="top" align="center">0.898</td>
</tr> <tr>
<td valign="top" align="center">Moisture content category (%)</td>
<td valign="top" align="center">1</td>
<td valign="top" align="center">64.036</td>
<td valign="top" align="center">149.415</td>
<td valign="top" align="center">&#x0003C; 0.001</td>
</tr> <tr style="background-color:#dee1e1">
<td valign="top" align="left" colspan="5"><bold>Tests of fixed effects (penetration resistance MPa)</bold></td>
</tr> <tr>
<td valign="top" align="center">Intercept</td>
<td valign="top" align="center">1</td>
<td valign="top" align="center">5.958</td>
<td valign="top" align="center">226.039</td>
<td valign="top" align="center">&#x0003C; 0.001</td>
</tr> <tr>
<td valign="top" align="center">Traction winch</td>
<td valign="top" align="center">1</td>
<td valign="top" align="center">11.217</td>
<td valign="top" align="center">2.222</td>
<td valign="top" align="center">0.164</td>
</tr> <tr>
<td valign="top" align="center">Slash cover</td>
<td valign="top" align="center">1</td>
<td valign="top" align="center">66.962</td>
<td valign="top" align="center">3.134</td>
<td valign="top" align="center">0.081</td>
</tr> <tr>
<td valign="top" align="center">Slope category (%)</td>
<td valign="top" align="center">4</td>
<td valign="top" align="center">50.229</td>
<td valign="top" align="center">0.913</td>
<td valign="top" align="center">0.464</td>
</tr> <tr>
<td valign="top" align="center">Moisture content category (%)</td>
<td valign="top" align="center">4</td>
<td valign="top" align="center">65.426</td>
<td valign="top" align="center">1.704</td>
<td valign="top" align="center">0.160</td>
</tr> <tr style="background-color:#dee1e1">
<td valign="top" align="left" colspan="5"><bold>Tests of fixed effects (maximal rut depth cm)</bold></td>
</tr> <tr>
<td valign="top" align="center">Intercept</td>
<td valign="top" align="center">1</td>
<td valign="top" align="center">3.850</td>
<td valign="top" align="center">181.248</td>
<td valign="top" align="center">&#x0003C; 0.001</td>
</tr> <tr>
<td valign="top" align="center">Traction winch</td>
<td valign="top" align="center">1</td>
<td valign="top" align="center">7.127</td>
<td valign="top" align="center">9.091</td>
<td valign="top" align="center">0.019</td>
</tr> <tr>
<td valign="top" align="center">Slash cover</td>
<td valign="top" align="center">1</td>
<td valign="top" align="center">66.672</td>
<td valign="top" align="center">1.231</td>
<td valign="top" align="center">0.271</td>
</tr> <tr>
<td valign="top" align="center">Slope category (%)</td>
<td valign="top" align="center">4</td>
<td valign="top" align="center">61.635</td>
<td valign="top" align="center">1.724</td>
<td valign="top" align="center">0.156</td>
</tr> <tr>
<td valign="top" align="center">Moisture content category (%)</td>
<td valign="top" align="center">4</td>
<td valign="top" align="center">65.944</td>
<td valign="top" align="center">0.379</td>
<td valign="top" align="center">0.823</td>
</tr></tbody>
</table>
</table-wrap>
</sec>
<sec>
<title>3.2 Penetration resistance and penetration depth</title>
<p>Penetration resistance increased in the rut location, from 4.88 (no. 1) to 26.8% (Control). <xref ref-type="table" rid="T6">Table 6</xref> shows that the increase in penetration resistances between the rut and stand locations was largest at the Control stand. Compared to the stand location (2.50&#x02013;3.10 MPa), the PR in the middle of the trails (2.57&#x02013;3.06 MPa) varied in some cases, it increased by 8 (Control) to 17 % (no. 2), while in other cases, it decreased by 1.29 (no. 3)&#x02212;10.45 % (no. 1). There was a decrease of penetration depth (cm) 1.96 (no. 3)&#x02212;10.95% (no. 2) in ruts (18.98&#x02013;19.77 cm) compared to the stand location (19.36&#x02013;22.20 cm) because the cone&#x00027;s ability to penetrate the soil was decreased as a result of machinery traffic. Interestingly, in stand no. 1, the penetration depth (cm) increased by 33.48% (4.8 cm) in the ruts compared to the stand location (<xref ref-type="table" rid="T6">Table 6</xref>). At the middle location, penetration depth (cm) increased by 0.26 (no. 3)&#x02212;17.83% (no. 1), compared to the stand location, except for stand no. 2, where we observed a decrease in penetration depth by 7.79%. One-way ANOVA (Duncan&#x00027;s test) confirmed that the differences between the forest stands (no. 1, 2, 3, Control) were significant on the stand (<italic>F</italic> = 19.29; <italic>p</italic> = 0.00), rut (<italic>F</italic> = 7.759; <italic>p</italic> = 0.00), and middle (<italic>F</italic>= 54.367; <italic>p</italic> = 0.00) measurement locations. The differences between the stand, rut, and middle measurement locations (Tukey&#x00027;s test) were also significant in all forest stands (<italic>p</italic> &#x0003C; 0.05). In the case of penetration depth (cm), Duncan&#x00027;s test confirmed that for the same measurement locations, the differences between forest stands (no. 1, 2, 3, Control) were significant (<italic>p</italic> &#x0003C; 0.05), except for the rut location (<italic>F</italic> = 1.368; <italic>p</italic> = 0.25). Tukey&#x00027;s test showed that the differences between the stand, rut, and middle measurement locations were significant in all stands except for stand no. 3 (<italic>F</italic> = 0.189; <italic>p</italic> = 0.828) (<xref ref-type="table" rid="T6">Table 6</xref>).</p>
<table-wrap position="float" id="T6">
<label>Table 6</label>
<caption><p>Mean penetration resistance and penetration depth values and their differences (%) compared through Tukey&#x00027;s test between measurement locations and observed stand no. (Duncan&#x00027;s test).</p></caption>
<table frame="box" rules="all">
<thead>
<tr style="background-color:#919498;color:#ffffff">
<th/>
<th valign="top" align="left" colspan="2"><bold>Stand (S)</bold></th>
<th valign="top" align="center" colspan="2"><bold>Rut (R)</bold></th>
<th valign="top" align="center" colspan="2"><bold>Middle (M)</bold></th>
<th/>
</tr>
</thead>
<tbody>
<tr style="background-color:#919498;color:#ffffff">
<td valign="top" align="center"><bold>Stand no</bold>.</td>
<td valign="top" align="left"><bold>&#x000D8; PR</bold><sup><italic>a</italic>)</sup> &#x000B1;<bold>SD</bold><sup><italic>b</italic>)</sup></td>
<td valign="top" align="center">&#x00394;<sup><italic>c</italic>)</sup> <bold>S</bold>&#x02192;<bold>R (%)</bold></td>
<td valign="top" align="center"><bold>&#x000D8; PR</bold><sup><italic>d</italic>)</sup> &#x000B1;<bold>SD</bold></td>
<td valign="top" align="center">&#x00394; <bold>R</bold>&#x02192;<bold>M (%)</bold></td>
<td valign="top" align="left"><bold>&#x000D8; PR</bold><sup><italic>d</italic>)</sup> &#x000B1;<bold>SD</bold></td>
<td valign="top" align="center">&#x00394; <bold>S</bold>&#x02192;<bold>M (%)</bold></td>
<td valign="top" align="center"><bold>Tukey</bold><sup><italic>e</italic>)</sup> &#x021C6;</td>
</tr> <tr>
<td valign="top" align="center">1</td>
<td valign="top" align="left">2.87 &#x000B1; 1.58</td>
<td valign="top" align="center">&#x0002B;4.88</td>
<td valign="top" align="center">3.01 &#x000B1; 1.50</td>
<td valign="top" align="center"><bold>&#x02212;14.62</bold></td>
<td valign="top" align="left">2.57 &#x000B1; 1.50</td>
<td valign="top" align="center"><bold>&#x02212;10.45</bold></td>
<td valign="top" align="center">R&#x02013;M; S&#x02013;M</td>
</tr> <tr>
<td valign="top" align="center">2</td>
<td valign="top" align="left">3 &#x000B1; 1.38</td>
<td valign="top" align="center"><bold>&#x0002B;13</bold></td>
<td valign="top" align="center">3.39 &#x000B1; 1.43</td>
<td valign="top" align="center">&#x0002B;3.54</td>
<td valign="top" align="left">3.51 &#x000B1; 1.70</td>
<td valign="top" align="center"><bold>&#x0002B;17</bold></td>
<td valign="top" align="center">S&#x02013;R; S&#x02013;M</td>
</tr> <tr>
<td valign="top" align="center">3</td>
<td valign="top" align="left">3.10 &#x000B1; 1.28</td>
<td valign="top" align="center"><bold>&#x0002B;7.42</bold></td>
<td valign="top" align="center">3.33 &#x000B1; 1.80</td>
<td valign="top" align="center"><bold>&#x02212;8.11</bold></td>
<td valign="top" align="left">3.06 &#x000B1; 1.68</td>
<td valign="top" align="center">&#x02212;1.29</td>
<td valign="top" align="center">S&#x02013;R; R&#x02013;M</td>
</tr> <tr>
<td valign="top" align="center">Control</td>
<td valign="top" align="left">2.50 &#x000B1; 0.96</td>
<td valign="top" align="center"><bold>&#x0002B;26.8</bold></td>
<td valign="top" align="center">3.17 &#x000B1; 1.79</td>
<td valign="top" align="center"><bold>&#x02212;14.83</bold></td>
<td valign="top" align="left">2.70 &#x000B1; 1.39</td>
<td valign="top" align="center">&#x0002B; 8</td>
<td valign="top" align="center">S&#x02013;R; R&#x02013;M</td>
</tr> <tr>
<td valign="top" align="center">Duncan<sup><italic>f</italic>)</sup> &#x02195;</td>
<td valign="top" align="left">1&#x02013;3; 1&#x02013;Con.; 2&#x02013;Con.; 3&#x02013;Con.</td>
<td valign="top" align="center"><bold>&#x02013;</bold></td>
<td valign="top" align="center">1&#x02013;2; 1&#x02013;3; 2&#x02013; Con.</td>
<td valign="top" align="center"><bold>&#x02013;</bold></td>
<td valign="top" align="left">1&#x02013;2; 1&#x02013;3; 2&#x02013;Con.; 2&#x02013;3; 3&#x02013; Con.</td>
<td valign="top" align="center"><bold>&#x02013;</bold></td>
<td valign="top" align="center">&#x02013;</td>
</tr> <tr style="background-color:#919498;color:#ffffff">
<td valign="top" align="center"><bold>Stand no</bold>.</td>
<td valign="top" align="left"><bold>&#x000D8; PD</bold> &#x000B1;<bold>SD</bold></td>
<td valign="top" align="center">&#x00394; <bold>S</bold>&#x02192;<bold>R (%)</bold></td>
<td valign="top" align="center"><bold>&#x000D8; PD</bold> &#x000B1;<bold>SD</bold></td>
<td valign="top" align="center">&#x00394;<sup><italic>d</italic>)</sup> <bold>R</bold>&#x02192;<bold>M (%)</bold></td>
<td valign="top" align="left"><bold>&#x000D8; PD</bold> &#x000B1;<bold>SD</bold></td>
<td valign="top" align="center">&#x00394; <bold>S</bold>&#x02192;<bold>M (%)</bold></td>
<td valign="top" align="center"><bold>Tukey</bold><sup><italic>e</italic>)</sup> &#x021C6;</td>
</tr> <tr>
<td valign="top" align="center">1</td>
<td valign="top" align="left">18.79 &#x000B1; 13.98</td>
<td valign="top" align="center"><bold>&#x0002B;33.48</bold></td>
<td valign="top" align="center">25.08 &#x000B1; 18.32</td>
<td valign="top" align="center"><bold>&#x02212;11.72</bold></td>
<td valign="top" align="left">22.14 &#x000B1; 15.72</td>
<td valign="top" align="center"><bold>&#x0002B;17.83</bold></td>
<td valign="top" align="center">S&#x02013;R; R&#x02013;M; S&#x02013;M</td>
</tr> <tr>
<td valign="top" align="center">2</td>
<td valign="top" align="left">22.20 &#x000B1; 18.27</td>
<td valign="top" align="center"><bold>&#x02212;10.95</bold></td>
<td valign="top" align="center">19.77 &#x000B1; 17.39</td>
<td valign="top" align="center">&#x0002B;3.54</td>
<td valign="top" align="left">20.47 &#x000B1; 15.38</td>
<td valign="top" align="center">&#x02212;7.79</td>
<td valign="top" align="center">S&#x02013;R</td>
</tr> <tr>
<td valign="top" align="center">3</td>
<td valign="top" align="left">19.36 &#x000B1; 13.19</td>
<td valign="top" align="center">&#x02212;1.96</td>
<td valign="top" align="center">18.98 &#x000B1; 14.11</td>
<td valign="top" align="center">&#x0002B;2.27</td>
<td valign="top" align="left">19.41 &#x000B1; 15.90</td>
<td valign="top" align="center">&#x0002B;0.26</td>
<td valign="top" align="center"><italic>p</italic> &#x0003E; 0.05</td>
</tr> <tr>
<td valign="top" align="center">Control</td>
<td valign="top" align="left">22.50 &#x000B1; 18.04</td>
<td valign="top" align="center">&#x02212;9.11</td>
<td valign="top" align="center">20.45 &#x000B1; 15.87</td>
<td valign="top" align="center"><bold>&#x0002B;14.57</bold></td>
<td valign="top" align="left">23.43 &#x000B1; 16.83</td>
<td valign="top" align="center">&#x0002B;4.13</td>
<td valign="top" align="center">R&#x02013;M</td>
</tr> <tr>
<td valign="top" align="center">Duncan<sup><italic>f</italic>)</sup> &#x02195;</td>
<td valign="top" align="left">1&#x02013;3; 2&#x02013;3; 3&#x02013;Con.</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center"><italic>p</italic> &#x0003E; 0.05</td>
<td valign="top" align="center"><bold>&#x02013;</bold></td>
<td valign="top" align="left">1&#x02013;2; 1&#x02013;3; 2&#x02013;Con.; 3&#x02013;Con.</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">&#x02013;</td>
</tr></tbody>
</table>
<table-wrap-foot>
<p><sup><italic>a</italic>)</sup>PR, penetration resistance; <sup><italic>b</italic>)</sup>SD, standard deviation; <sup><italic>c</italic>)</sup>&#x00394;, value difference; bold typeface indicates the significance of differences (<italic>p</italic> &#x02264; 0.05) between locations; <sup><italic>d</italic>)</sup>PD, penetration depth; <sup><italic>e</italic>)</sup>Tukey, between-group comparisons of differences between the measurement locations within each particular forest stand; <sup><italic>f</italic>)</sup>Duncan, between-group comparisons of differences of the same measurement location between particular forest stands.</p>
</table-wrap-foot>
</table-wrap>
<p>Linear mixed modeling between PR as the dependent variable, measurement location as fixed effects, and stand no. as random effects showed that only 0.7% (<inline-formula><mml:math id="M6"><mml:msubsup><mml:mrow><mml:mi>R</mml:mi></mml:mrow><mml:mrow><mml:mtext>m</mml:mtext></mml:mrow><mml:mrow><mml:mn>2</mml:mn></mml:mrow></mml:msubsup></mml:math></inline-formula> = 0.007) of PR variability were affected by measurement location, and further 4.3% were affected by fixed and random effects (<italic>R</italic><sup>2</sup>c = 0.05) and the relationship was insignificant (<italic>F</italic> = 0.446; <italic>p</italic> = 0.64). We achieved similar results when penetration depth (cm) was the dependent variable (<italic>R</italic><sup>2</sup>m = 0.005, <italic>R</italic><sup>2</sup>c = 0.012). The LMM between the rut PR, stand no. (random effect), and the usage of traction winch (<italic>F</italic> = 2.222, <italic>p</italic> = 0.164), presence of slash in the ruts (<italic>F</italic> = 3.134, <italic>p</italic> = 0.081), slope category (<italic>F</italic> = 0.913, <italic>p</italic> = 0.464), and moisture content category (<italic>F</italic> = 1.704, <italic>p</italic> = 0.160) (fixed effects) showed no significant relationship (<xref ref-type="table" rid="T5">Table 5</xref>). Another LMM did not confirm a significant relationship between PR and BD (<italic>F</italic> = 0.901; <italic>p</italic> = 0.346; <italic>R</italic><sup>2</sup>m = 0.012).</p>
</sec>
<sec>
<title>3.3 Slash and rut depth</title>
<p>Slash (treetops and limbs) were placed on the trails by the operator during the harvesting process to minimize the effects of the machine on the soil surface. From measured data, it can be seen that the share of measurements without slash placed on the trails was relatively high, ranging from 67 to 95% (<xref ref-type="table" rid="T7">Table 7</xref>). Consequently, LMM in the ruts of the forwarding trails did not confirm a significant relationship (<italic>F</italic> = 0.264; <italic>p</italic> = 0.769) between the slash thickness category (cm) and BD (<italic>R</italic><sup>2</sup>m = 0.007; <italic>R</italic><sup>2</sup>c = 0.094). In the case of PR and slash thickness category (cm), the relationship was significant (<italic>F</italic> = 3.819; <italic>p</italic> = 0.027), though the explanatory power was weak <italic>R</italic><sup>2</sup>m (0.090) and <italic>R</italic><sup>2</sup>c (0.094). Insignificant or weak relationships between slash cover and BD or PR were caused by the small amount of slash cover over the ruts of the forwarding trails, which ranged between 4.17 and 29.16%. An inadequate layer of slash on the trail or its absence was mainly caused by the harvesting operations being carried out in young, 30- to 40-year-old stands. The thin branches and the low harvesting concentrations did not permit covering the trails with sufficient slash.</p>
<table-wrap position="float" id="T7">
<label>Table 7</label>
<caption><p>The cover of trails with slash and its thickness in 10 cm intervals recorded at particular measurement locations.</p></caption>
<table frame="box" rules="all">
<thead>
<tr style="background-color:#919498;color:#ffffff">
<th/>
<th valign="top" align="center" colspan="2"><bold>1</bold></th>
<th valign="top" align="center" colspan="2"><bold>2</bold></th>
<th valign="top" align="center" colspan="2"><bold>3</bold></th>
<th valign="top" align="center" colspan="2"><bold>Control</bold></th>
</tr>
</thead>
<tbody>
<tr style="background-color:#919498;color:#ffffff">
<td/>
<td valign="top" align="center"><bold>Rut</bold></td>
<td valign="top" align="center"><bold>Middle</bold></td>
<td valign="top" align="center"><bold>Rut</bold></td>
<td valign="top" align="center"><bold>Middle</bold></td>
<td valign="top" align="center"><bold>Rut</bold></td>
<td valign="top" align="center"><bold>Middle</bold></td>
<td valign="top" align="center"><bold>Rut</bold></td>
<td valign="top" align="center"><bold>Middle</bold></td>
</tr> <tr>
<td valign="top" align="center">No slash (<italic>n</italic>/%)<sup>a</sup></td>
<td valign="top" align="center">16/66.67</td>
<td valign="top" align="center">3/12.5</td>
<td valign="top" align="center">19/95</td>
<td valign="top" align="center">14/70</td>
<td valign="top" align="center">15/68.18</td>
<td valign="top" align="center">6/27.27</td>
<td valign="top" align="center">10/83.33</td>
<td valign="top" align="center">5/41.67</td>
</tr> <tr>
<td valign="top" align="center">1&#x02013;10 cm (<italic>n</italic>/%)</td>
<td valign="top" align="center">7/29.16</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">5/22.73</td>
<td valign="top" align="center">1/4.54</td>
<td valign="top" align="center">2/16.67</td>
<td valign="top" align="center">&#x02013;</td>
</tr> <tr>
<td valign="top" align="center">11&#x02013;20 cm (<italic>n</italic>/%)</td>
<td valign="top" align="center">1/4.17</td>
<td valign="top" align="center">7/29.17</td>
<td valign="top" align="center">1/5</td>
<td valign="top" align="center">2/10</td>
<td valign="top" align="center">2/9.09</td>
<td valign="top" align="center">8/36.37</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">4/33.33</td>
</tr> <tr>
<td valign="top" align="center">21&#x02013;30 cm (<italic>n</italic>/%)</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">12/50</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">2/10</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">7/31.82</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">2/16.67</td>
</tr> <tr>
<td valign="top" align="center">31&#x02013;40 cm (<italic>n</italic>/%)</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">2/8.33</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">2/10</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">&#x02013;</td>
<td valign="top" align="center">1/8.33</td>
</tr> <tr>
<td valign="top" align="center">&#x02211;</td>
<td valign="top" align="center">24/100</td>
<td valign="top" align="center">24/100</td>
<td valign="top" align="center">20/100</td>
<td valign="top" align="center">20/100</td>
<td valign="top" align="center">22/100</td>
<td valign="top" align="center">22/100</td>
<td valign="top" align="center">12/100</td>
<td valign="top" align="center">12/100</td>
</tr></tbody>
</table>
<table-wrap-foot>
<p><sup><italic>a</italic>)</sup><italic>n</italic>/%, number of measurement places in a stand/percentage share.</p>
</table-wrap-foot>
</table-wrap>
<p>Mean forwarding trail profile depth in particular forest stands varied between 4.63 and 7.28 cm, whereas maximum rut depth ranged between 10.86 and 17.25 cm (<xref ref-type="table" rid="T8">Table 8</xref>). The deepest average trail profile and maximum rut depth were recorded in the Control stand.</p>
<table-wrap position="float" id="T8">
<label>Table 8</label>
<caption><p>Mean and maximum rut depth and percentage share of trail disturbance classes in particular forest stands.</p></caption>
<table frame="box" rules="all">
<thead>
<tr style="background-color:#919498;color:#ffffff">
<th valign="top" align="left"><bold>Stand</bold></th>
<th valign="top" align="center"><bold>1</bold>.</th>
<th valign="top" align="center"><bold>2</bold>.</th>
<th valign="top" align="center"><bold>3</bold>.</th>
<th valign="top" align="center"><bold>Control</bold></th>
</tr>
</thead>
<tbody>
<tr>
<td valign="top" align="left">Mean rut depth &#x000B1; SD (cm)</td>
<td valign="top" align="center">5.08 &#x000B1; 1.1</td>
<td valign="top" align="center">5.59 &#x000B1; 1.1</td>
<td valign="top" align="center">4.63 &#x000B1; 1.4</td>
<td valign="top" align="center">7.28 &#x000B1; 3.0</td>
</tr> <tr>
<td valign="top" align="left">Maximum rut depth &#x000B1; SD (cm)</td>
<td valign="top" align="center">12.92 &#x000B1; 3.2</td>
<td valign="top" align="center">13 &#x000B1; 3.1</td>
<td valign="top" align="center">10.86 &#x000B1; 3.3</td>
<td valign="top" align="center">17.25 &#x000B1; 5.9</td>
</tr> <tr>
<td valign="top" align="left">Low disturbance &#x02264; 7.9 cm (%)</td>
<td valign="top" align="center">4.16</td>
<td valign="top" align="center">0</td>
<td valign="top" align="center">13.64</td>
<td valign="top" align="center">0</td>
</tr> <tr>
<td valign="top" align="left">Moderate disturbance 8&#x02013;15.9 cm (%)</td>
<td valign="top" align="center">79.17</td>
<td valign="top" align="center">80</td>
<td valign="top" align="center">77.27</td>
<td valign="top" align="center">41.67</td>
</tr> <tr>
<td valign="top" align="left">Severe disturbance 16&#x02013;25.9 cm (%)</td>
<td valign="top" align="center">16.67</td>
<td valign="top" align="center">20</td>
<td valign="top" align="center">9.09</td>
<td valign="top" align="center">50</td>
</tr> <tr>
<td valign="top" align="left">Very severe disturbance &#x02265; 26 cm (%)</td>
<td valign="top" align="center">0</td>
<td valign="top" align="center">0</td>
<td valign="top" align="center">0</td>
<td valign="top" align="center">8.33</td>
</tr></tbody>
</table>
<table-wrap-foot>
<p><sup><italic>a</italic>)</sup>SD, standard deviation.</p>
</table-wrap-foot>
</table-wrap>
<p>The most frequent intensity class of forwarding trail disturbance was a moderately strong disturbance, with a maximum rut depth between 8 and 15.9 cm (<xref ref-type="table" rid="T8">Table 8</xref>). This level of disturbance represented 77.27 (stand no. 3) to 80% (stand no. 2) of the trails in the stands. In the Control stand, severe disturbance was the most frequent, representing 50 % of disturbances, with a maximum rut depth between 16 and 25.9 cm. One-way ANOVA (Duncan&#x00027;s test) confirmed significant differences between stands. In the case of average rut depth (cm) (<italic>F</italic> = 7.21; <italic>p</italic> = 0.00) and maximum rut depth (<italic>F</italic>= 7.48; <italic>p</italic> = 0.00), the differences were significant in the Control stand, which differed significantly from all other forest stands (nos. 1, 2, 3).</p>
<p>The LMM for maximum rut depth (dependent variable), traction winch usage, slash cover, slope category, and moisture content as fixed effects, and stand no. as a random effect showed a significant relationship between the traction winch usage (<italic>F</italic> = 9.091; <italic>p</italic> = 0.019) (<xref ref-type="table" rid="T5">Table 5</xref>). In the case of other variables, i.e., slash cover (<italic>F</italic> = 1.231; <italic>p</italic> = 0.271), slope category (<italic>F</italic> = 1.724; <italic>p</italic> = 0.156), and moisture content category (<italic>F</italic> = 0.379; <italic>p</italic> = 0.823), the relationships were insignificant.</p></sec>
</sec>
<sec id="s4">
<title>4 Discussion</title>
<p>Soil disturbance caused by deploying harvesters assisted by mobile traction or other winches in thinnings is not the subject of many studies. Multiple studies research the safety, tensile forces, slope range, efficiency, and productivity of this technology on steep slopes (Holzfeind et al., <xref ref-type="bibr" rid="B14">2018</xref>; Holzleitner et al., <xref ref-type="bibr" rid="B15">2018</xref>; Jod&#x00142;owski and Kalinowski, <xref ref-type="bibr" rid="B18">2018</xref>; Mologni et al., <xref ref-type="bibr" rid="B28">2018</xref>, <xref ref-type="bibr" rid="B29">2021</xref>; Visser and Stampfer, <xref ref-type="bibr" rid="B42">2015</xref>). Literature also states that using winches to secure ground-based harvesting machines can contribute to soil disturbance reduction due to the ability of the tethered system to avoid sliding, slipping, and digging farther into the soil (Cavalli and Amishev, <xref ref-type="bibr" rid="B7">2019</xref>; Holzleitner et al., <xref ref-type="bibr" rid="B15">2018</xref>). The winch can support the braking and traction of the harvesting machine, thus distributing the forces and minimizing the wheel slippage of the machine when navigating challenging terrains. Severe soil disturbance is reduced because wheel slippage decreases to &#x0003C; 20%, making it easier to develop logging trails systematically. However, these are theoretical statements without direct measurements of the effects of this technology on soil disturbance.</p>
<sec>
<title>4.1 Bulk density</title>
<p>Garren et al. (<xref ref-type="bibr" rid="B10">2019</xref>) studied the changes in soil parameters when mobile traction winches are used in steep slopes (27&#x02013;38&#x000B0;, i.e., 51&#x02013;78%), dividing the slopes into three categories (first: 27&#x02013;31&#x000B0;, i.e., 51&#x02013;60%; second: 31&#x02013;35&#x000B0;, i.e., 60&#x02013;70%; third: 35&#x02013;38&#x000B0;, i.e., 70&#x02013;78%). To compare, we chose the first slope category from the study by Garren et al. (<xref ref-type="bibr" rid="B10">2019</xref>). In that category, the bulk density in rut ranged from 1.4 to 1.58 g cm<sup>&#x02212;3</sup> and increased by 16% on average compared to stand locations. In our case, at 22&#x02013;29.5&#x000B0; (40.4&#x02013;56.6%) slopes, the bulk density in rut ranged between 0.93 and 1.05 g cm<sup>&#x02212;3</sup> and increased by 13.42 (stand no. 2)&#x02212;22.09% (stand no. 3). At lower slopes (14.9&#x000B0; i.e., 26.6% stand Control), higher bulk density was not observed in the rut (1.04 g cm<sup>&#x02212;3</sup>) compared to other stands, though the increase was more substantial (22.35%) than in stands with higher slopes. The LMM did not confirm a statistically significant effect of slope on the BD (<italic>p</italic> &#x0003E; 0.05). To our knowledge, very few studies focus on the problem of using mobile traction winches combined with CTL machinery in thinnings and the soil disturbance caused by this harvesting system. However, we can use studies of non-assisted CTL machinery for comparisons. Malvar et al. (<xref ref-type="bibr" rid="B26">2017</xref>) indicate that mean BD at 0&#x02013;5 cm depth ranges from 0.77 kg cm<sup>&#x02212;3</sup> at the stand area to 1.07 kg cm<sup>&#x02212;3</sup> at compacted rut area, and BD is between 14 and 28% higher in rut. This research was conducted on (47&#x02013;53%, i.e., approximately 25.2&#x02013;27.9&#x000B0;) slope. Allman et al. (<xref ref-type="bibr" rid="B1">2015</xref>) report an increase in soil bulk density from (0.82 to 1.12 g cm<sup>&#x02212;3</sup>) in stand to (1.02 to 1.36 g cm<sup>&#x02212;3</sup>) in rut, i.e., between 21 and 2% after CTL machine traffic. Labelle et al. (<xref ref-type="bibr" rid="B20">2019</xref>) indicate a pre-operation soil BD from 0.4 to 1.0 g&#x000B7;cm<sup>&#x02212;3</sup> and from 0.7 to 1.2 g cm<sup>&#x02212;3</sup> after machine traffic, representing an increase between 20 and 75%. Williamson and Neilsen (<xref ref-type="bibr" rid="B44">2000</xref>) report a BD increase of 0.17 g.cm<sup>&#x02212;3</sup> after machine traffic.</p>
</sec>
<sec>
<title>4.2 Penetration resistance</title>
<p>Penetration resistance increased by 4.8&#x02013;26.8% in the rut, while the penetration depth decreased by 2&#x02013;10%. The most substantial increase in PR (26.8%) was observed in stand Control, where the winch did not operate. However, according to Tukey&#x00027;s test, the differences between the stand and rut locations were significant in all forest stands. Labelle et al. (<xref ref-type="bibr" rid="B20">2019</xref>) state that on forest or agricultural land, high soil PR may be problematic for root growth and values beyond 3.0 MPa are regarded as a threshold for interrupted root growth, thus marking severely disturbed soils. In our case, PR over 3.0 MPa was recorded even in undisturbed locations, although only in the case of stand no. 3. To compare, PR values in the rut ranged between 3.01 and 3.17 MPa. Labelle et al. (<xref ref-type="bibr" rid="B20">2019</xref>) concluded that while none of the pre-forwarding measurement points exceeded the 3.0 MPa threshold, 10.7% of post-forwarding measurement points showed PR above 3.0 MPa, with about 80% of such values occurring in the rut. We presume that the higher PR values in the stand locations in our case were due to insufficient rainfall during August and September, as well as moisture deficit in lower soil layers. On the other hand, the LMM did not confirm a statistically significant relationship between the soil moisture content on the PR (<italic>p</italic> &#x0003E; 0.05), likely due to our use of determining the moisture content from samples taken from upper soil layers via a gravimetric method. In contrast, the PR values were recorded at depths up to 80 cm, where different moisture conditions can occur, which might have caused an increased PR when trying to penetrate the cone penetrometer into deeper soil layers. Cudzik et al. (<xref ref-type="bibr" rid="B8">2017</xref>) report even higher PR values. Their study showed that PR after CTL machine traffic increased by 220&#x02013;308% compared to control measurements at the stand location. The penetration resistances measured ranged from 2.8 to even 4 MPa.</p>
</sec>
<sec>
<title>4.3 Rut depth and soil moisture</title>
<p>In their study, Garren et al. (<xref ref-type="bibr" rid="B10">2019</xref>) indicate that the maximum rut depth (cm) caused by the winch technology was 9.5 cm and in 39% of observations, the rut depth was 0 cm or no visible ruts and overall rutting was negligible. In our case, the maximum rut depth ranged between 10.86 and 17.25 cm. In more than one-quarter to one-fifth of observations where winch assistance was used, the soil disturbance reached a &#x0201C;moderately strong&#x0201D; category, while in the stand without a winch, the rutting reached a &#x0201C;severe disturbance&#x0201D; level most frequently, though the differences in rut depth (cm) were primarily insignificant. Visser and Spinelli (<xref ref-type="bibr" rid="B43">2023</xref>) state that on steeper slopes (30&#x02013;40%, i.e., 16.7&#x02013;21.8&#x000B0;), low soil disturbance and rut deeper than 15 cm covers only 2% of the observed surface after being trafficked by a two-axle skidder with winch assistance. In our case, ruts deeper than 15 cm were measured frequently, between 9 (stand no. 3) and 50% (Control) of measurement locations. Although traction-assist technology significantly reduces wheel slippage, no mitigating effect of traction-assist technology on soil disturbance was observed. Cudzik et al. (<xref ref-type="bibr" rid="B8">2017</xref>) report a rut depth of up to 5 cm on 81% of trails in their study of CTL machines working in slopes between 8 and 12&#x000B0; (14&#x02013;21.3%). Han et al. (<xref ref-type="bibr" rid="B13">2006</xref>) state that ruts are deeper for moist (medium and high moisture contents) than for drier soils. The high moisture content (%) contributes to the development of ruts that are significantly (13.8&#x02013;35.3 cm) deeper than in the case of medium (8.2&#x02013;12.0 cm) or low moisture contents (4.5&#x02013;8.0 cm). In our case, the deepest ruts were recorded in the forest stand, where the CTL machines operated unassisted, as confirmed by the fact that the LMM showed a statistically significant effect of a traction winch being used (<italic>p</italic> &#x0003C; 0.05) on the maximum rut depth in the forwarding trails. Duncan&#x00027;s test confirmed significant differences between the mean and maximum rut depth between the Control stand and stands no. 1, 2, and 3.</p>
</sec>
<sec>
<title>4.4 Study limitations</title>
<p>Forest harvesting in mountainous regions of the Carpathian mountain range happens in various technological conditions connected to machinery and their use, soil types, terrain configuration, and tree species being harvested. Therefore, large-scale studies are needed to cover the array of machines and their use in highly variable natural conditions. Future research should expand the use cases of traction winches supporting ground-based wheeled machinery and focus on repeated observations with control plots that reflect the variability of natural conditions. Different approaches to observing the soil disturbance complementary to the methods used in this study can also be used in the future.</p>
<p>Admittedly, our study has a limited scale. We could only perform control measurements in one comparable forest stand, while more would be preferable. Our inability to do so was caused by the fact that no suitable forest stands were being harvested by the land owner where the slope would be sufficient on the one hand, and occupational safety and health could be secured on the other since the operation of unassisted machinery in slopes over 40% slope was forbidden by the internal regulations of the land owner.</p>
<p>Nevertheless, even small-scale studies contribute to filling in the considerable knowledge gap on the environmental effects of this technological variant and providing practitioners with adequate information when considering what technology and machinery to deploy in their forests.</p></sec>
</sec>
<sec id="s5">
<title>5 Conclusion</title>
<p>Based on our findings, we reached the following conclusions:</p>
<list list-type="bullet">
<list-item><p>In the case of BD, the winch-assisted machines did not cause a higher degree of compaction compared to unassisted machines. The LMM did not confirm a significant effect of traction winches on the BD (<italic>p</italic> &#x0003E; 0.05). Hypothesis H<sub>1</sub> was therefore rejected, and we can state that the two technology variants caused a similar degree of soil compaction.</p></list-item>
<list-item><p>The most substantial increase in soil penetration resistance in the ruts compared to control locations (stand) was recorded in the control stand. A significant PR increase compared to the Control stand was confirmed in the case of stands no. 2 and 3, while the LMM did not confirm a significant effect of the traction winch on PR in the rut of the forwarding trails (<italic>p</italic> &#x0003E; 0.05). Hypothesis H<sub>2</sub> was therefore rejected.</p></list-item>
<list-item><p>The deepest ruts and most substantial mean forwarding trail profile depth were observed in the stand where CTL machines worked unassisted by traction winches. On the other hand, significant differences (Duncan&#x00027;s test) were confirmed between Control and stands no. 1, 2, and 3. The LMM confirmed a significant effect of traction winch on the maximum rut depth. Hypothesis H<sub>3</sub> was, therefore, confirmed.</p></list-item>
</list>
<p>Management recommendations and implications:</p>
<list list-type="bullet">
<list-item><p>The use of winch-assisted technologies in steep slopes did not cause additional soil compaction or increased penetration resistance compared to those slopes where CTL machinery can work unassisted.</p></list-item>
<list-item><p>Using a traction winch causes a significantly smaller disturbance of the soil surface, as represented by the measurable depth of the ruts of the forwarding trails.</p></list-item>
<list-item><p>Mobile traction winches are a viable technological adaptation that enables deploying ground-based harvesting in terrains where cable yarders would otherwise have to operate.</p></list-item>
</list></sec>
</body>
<back>
<sec sec-type="data-availability" id="s6">
<title>Data availability statement</title>
<p>The data that support the findings of this study are available from the corresponding author upon reasonable request.</p>
</sec>
<sec sec-type="author-contributions" id="s7">
<title>Author contributions</title>
<p>MA: Writing &#x02013; original draft, Writing &#x02013; review &#x00026; editing, Conceptualization, Data curation, Methodology, Software, Visualization. ZD: Writing &#x02013; original draft, Writing &#x02013; review &#x00026; editing, Conceptualization, Formal analysis, Methodology, Visualization. MD: Writing &#x02013; original draft, Writing &#x02013; review &#x00026; editing, Methodology, Software, Validation, Visualization. MJ: Writing &#x02013; original draft, Writing &#x02013; review &#x00026; editing, Conceptualization, Data curation, Formal analysis, Methodology. VJ: Writing &#x02013; original draft, Writing &#x02013; review &#x00026; editing.</p>
</sec>
<sec sec-type="funding-information" id="s8">
<title>Funding</title>
<p>The author(s) declare financial support was received for the research, authorship, and/or publication of this article. Grant No. NAZV QK22020146 Guidelines for Water Management on Forest Transportation Networks supported this work.</p>
</sec>
<ack><p>We thank the Military forests and estates of the Slovak Republic, GOE, for enabling the measurements and supplying all data necessary for this study.</p>
</ack>
<sec sec-type="COI-statement" id="conf1">
<title>Conflict of interest</title>
<p>The authors declare that the research was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest.</p>
</sec>
<sec sec-type="disclaimer" id="s9">
<title>Publisher&#x00027;s note</title>
<p>All claims expressed in this article are solely those of the authors and do not necessarily represent those of their affiliated organizations, or those of the publisher, the editors and the reviewers. Any product that may be evaluated in this article, or claim that may be made by its manufacturer, is not guaranteed or endorsed by the publisher.</p>
</sec>
<sec sec-type="supplementary-material" id="s10">
<title>Supplementary material</title>
<p>The Supplementary Material for this article can be found online at: <ext-link ext-link-type="uri" xlink:href="https://www.frontiersin.org/articles/10.3389/ffgc.2024.1464140/full#supplementary-material">https://www.frontiersin.org/articles/10.3389/ffgc.2024.1464140/full#supplementary-material</ext-link></p>
<supplementary-material xlink:href="Image_1.pdf" id="SM1" mimetype="application/pdf" xmlns:xlink="http://www.w3.org/1999/xlink"/></sec>
<ref-list>
<title>References</title>
<ref id="B1">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Allman</surname> <given-names>M.</given-names></name> <name><surname>Jankovsk&#x000FD;</surname> <given-names>M.</given-names></name> <name><surname>Messingerov&#x000E1;</surname> <given-names>V.</given-names></name> <name><surname>Allmanov&#x000E1;</surname> <given-names>Z.</given-names></name> <name><surname>Feren&#x0010D;&#x000ED;k</surname> <given-names>M.</given-names></name></person-group> (<year>2015</year>). <article-title>Soil compaction of various Central European forest soils caused by traffic of forestry machines with various chassis</article-title>. <source>For. Syst.</source> <volume>24</volume>, <fpage>e038</fpage>&#x02013;<lpage>e038</lpage>. <pub-id pub-id-type="doi">10.5424/fs/2015243-07541</pub-id></citation>
</ref>
<ref id="B2">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Ares</surname> <given-names>A.</given-names></name> <name><surname>Terry</surname> <given-names>T. A.</given-names></name> <name><surname>Miller</surname> <given-names>R. E.</given-names></name> <name><surname>Anderson</surname> <given-names>H. W.</given-names></name> <name><surname>Flaming</surname> <given-names>B. L.</given-names></name></person-group> (<year>2005</year>). <article-title>Ground-based forest harvesting effects on soil physical properties and douglas-fir growth</article-title>. <source>Soil Sci. Soc. Am. J.</source> <volume>69</volume>, <fpage>1822</fpage>&#x02013;<lpage>1832</lpage>. <pub-id pub-id-type="doi">10.2136/sssaj2004.0331</pub-id></citation>
</ref>
<ref id="B3">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Blouin</surname> <given-names>V. M.</given-names></name> <name><surname>Schmidt</surname> <given-names>M. G.</given-names></name> <name><surname>Bulmer</surname> <given-names>C. E.</given-names></name> <name><surname>Krzic</surname> <given-names>M.</given-names></name></person-group> (<year>2008</year>). <article-title>Effects of compaction and water content on lodgepole pine seedling growth</article-title>. <source>For. Ecol. Manag.</source> <volume>255</volume>, <fpage>2444</fpage>&#x02013;<lpage>2452</lpage>. <pub-id pub-id-type="doi">10.1016/j.foreco.2008.01.008</pub-id></citation>
</ref>
<ref id="B4">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Cambi</surname> <given-names>M.</given-names></name> <name><surname>Certini</surname> <given-names>G.</given-names></name> <name><surname>Neri</surname> <given-names>F.</given-names></name> <name><surname>Marchi</surname> <given-names>E.</given-names></name></person-group> (<year>2015</year>). <article-title>The impact of heavy traffic on forest soils: a review</article-title>. <source>For. Ecol. Manag.</source> <volume>338</volume>, <fpage>124</fpage>&#x02013;<lpage>138</lpage>. <pub-id pub-id-type="doi">10.1016/j.foreco.2014.11.022</pub-id></citation>
</ref>
<ref id="B5">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Cambi</surname> <given-names>M.</given-names></name> <name><surname>Hoshika</surname> <given-names>Y.</given-names></name> <name><surname>Mariotti</surname> <given-names>B.</given-names></name> <name><surname>Paoletti</surname> <given-names>E.</given-names></name> <name><surname>Picchio</surname> <given-names>R.</given-names></name> <name><surname>Venanzi</surname> <given-names>R.</given-names></name> <etal/></person-group>. (<year>2017</year>). <article-title>Compaction by a forest machine affects soil quality and <italic>Quercus robur</italic> L. seedling performance in an experimental field</article-title>. <source>For. Ecol. Manag.</source> <volume>384</volume>, <fpage>406</fpage>&#x02013;<lpage>414</lpage>. <pub-id pub-id-type="doi">10.1016/j.foreco.2016.10.045</pub-id></citation>
</ref>
<ref id="B6">
<citation citation-type="web"><person-group person-group-type="author"><name><surname>Cavalli</surname> <given-names>R.</given-names></name></person-group> (<year>2012</year>). <article-title>Prospects of research on cable logging</article-title>. <source>Croat. J. Eng.</source> <volume>33</volume>, <fpage>339</fpage>&#x02013;<lpage>356</lpage>. Available at: <ext-link ext-link-type="uri" xlink:href="https://hrcak.srce.hr/116850">https://hrcak.srce.hr/116850</ext-link></citation>
</ref>
<ref id="B7">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Cavalli</surname> <given-names>R.</given-names></name> <name><surname>Amishev</surname> <given-names>D.</given-names></name></person-group> (<year>2019</year>). <article-title>Steep terrain forest operations &#x02013; challenges, technology development, current implementation, and future opportunities</article-title>. <source>Int. J. For. Eng.</source> <volume>30</volume>, <fpage>175</fpage>&#x02013;<lpage>181</lpage>. <pub-id pub-id-type="doi">10.1080/14942119.2019.1603030</pub-id></citation>
</ref>
<ref id="B8">
<citation citation-type="web"><person-group person-group-type="author"><name><surname>Cudzik</surname> <given-names>A.</given-names></name> <name><surname>Brennensthul</surname> <given-names>M.</given-names></name> <name><surname>Bia&#x00142;czyk</surname> <given-names>W.</given-names></name> <name><surname>Czarnecki</surname> <given-names>J.</given-names></name></person-group> (<year>2017</year>). <article-title>Damage to soil and residual trees caused</article-title>. <source>Croat. J. Eng</source>. <volume>38</volume>, <fpage>83</fpage>&#x02013;<lpage>95</lpage>. Available at: <ext-link ext-link-type="uri" xlink:href="https://hrcak.srce.hr/174453">https://hrcak.srce.hr/174453</ext-link></citation>
</ref>
<ref id="B9">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Etehadi Abari</surname> <given-names>M.</given-names></name> <name><surname>Majnounian</surname> <given-names>B.</given-names></name> <name><surname>Malekian</surname> <given-names>A.</given-names></name> <name><surname>Jourgholami</surname> <given-names>M.</given-names></name></person-group> (<year>2017</year>). <article-title>Effects of forest harvesting on runoff and sediment characteristics in the Hyrcanian forests, northern Iran</article-title>. <source>Eur. J. For. Res.</source> <volume>136</volume>, <fpage>375</fpage>&#x02013;<lpage>386</lpage>. <pub-id pub-id-type="doi">10.1007/s10342-017-1038-3</pub-id></citation>
</ref>
<ref id="B10">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Garren</surname> <given-names>A. M.</given-names></name> <name><surname>Bolding</surname> <given-names>M. C.</given-names></name> <name><surname>Aust</surname> <given-names>W. M.</given-names></name> <name><surname>Moura</surname> <given-names>A. C.</given-names></name> <name><surname>Barrett</surname> <given-names>S. M.</given-names></name></person-group> (<year>2019</year>). <article-title>Soil Disturbance effects from tethered forwarding on steep slopes in Brazilian eucalyptus plantations</article-title>. <source>Forests</source> <volume>10</volume>:<fpage>721</fpage>. <pub-id pub-id-type="doi">10.3390/f10090721</pub-id></citation>
</ref>
<ref id="B11">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Goutal</surname> <given-names>N.</given-names></name> <name><surname>Keller</surname> <given-names>T.</given-names></name> <name><surname>D&#x000E9;fossez</surname> <given-names>P.</given-names></name> <name><surname>Ranger</surname> <given-names>J.</given-names></name></person-group> (<year>2013</year>). <article-title>Soil compaction due to heavy forest traffic: measurements and simulations using an analytical soil compaction model</article-title>. <source>Ann. For. Sci.</source> <volume>70</volume>, <fpage>545</fpage>&#x02013;<lpage>556</lpage>. <pub-id pub-id-type="doi">10.1007/s13595-013-0276-x</pub-id></citation>
</ref>
<ref id="B12">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Green</surname> <given-names>P. Q.</given-names></name> <name><surname>Chung</surname> <given-names>W.</given-names></name> <name><surname>Leshchinsky</surname> <given-names>B.</given-names></name> <name><surname>Belart</surname> <given-names>F.</given-names></name> <name><surname>Sessions</surname> <given-names>J.</given-names></name> <name><surname>Fitzgerald</surname> <given-names>S. A.</given-names></name> <etal/></person-group>. (<year>2020</year>). <article-title>Insight into the productivity, cost and soil impacts of cable-assisted harvester-forwarder thinning in Western Oregon</article-title>. <source>For. Sci.</source> <volume>66</volume>, <fpage>82</fpage>&#x02013;<lpage>96</lpage>. <pub-id pub-id-type="doi">10.1093/forsci/fxz049</pub-id></citation>
</ref>
<ref id="B13">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Han</surname> <given-names>H.-S.</given-names></name> <name><surname>Page-Dumroese</surname> <given-names>D.</given-names></name> <name><surname>Han</surname> <given-names>S.-K.</given-names></name> <name><surname>Tirocke</surname> <given-names>J.</given-names></name></person-group> (<year>2006</year>). <article-title>Effects of slash, machine passes, and soil moisture on penetration resistance in a cut-to-length harvesting</article-title>. <source>Int. J. For. Eng.</source> <volume>17</volume>, <fpage>11</fpage>&#x02013;<lpage>24</lpage>. <pub-id pub-id-type="doi">10.1080/14942119.2006.10702532</pub-id></citation>
</ref>
<ref id="B14">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Holzfeind</surname> <given-names>T.</given-names></name> <name><surname>Stampfer</surname> <given-names>K.</given-names></name> <name><surname>Holzleitner</surname> <given-names>F.</given-names></name></person-group> (<year>2018</year>). <article-title>Productivity, setup time and costs of a winch-assisted forwarder</article-title>. <source>J. For. Res.</source> <volume>23</volume>, <fpage>196</fpage>&#x02013;<lpage>203</lpage>. <pub-id pub-id-type="doi">10.1080/13416979.2018.1483131</pub-id></citation>
</ref>
<ref id="B15">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Holzleitner</surname> <given-names>F.</given-names></name> <name><surname>Kastner</surname> <given-names>M.</given-names></name> <name><surname>Stampfer</surname> <given-names>K.</given-names></name> <name><surname>H&#x000F6;ller</surname> <given-names>N.</given-names></name> <name><surname>Kanzian</surname> <given-names>C.</given-names></name></person-group> (<year>2018</year>). <article-title>Monitoring cable tensile forces of winch-assist harvester and forwarder operations in steep terrain</article-title>. <source>Forests</source> <volume>9</volume>:<fpage>53</fpage>. <pub-id pub-id-type="doi">10.3390/f9020053</pub-id></citation>
</ref>
<ref id="B16">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Horn</surname> <given-names>R.</given-names></name> <name><surname>Vossbrink</surname> <given-names>J.</given-names></name> <name><surname>Becker</surname> <given-names>S.</given-names></name></person-group> (<year>2004</year>). <article-title>Modern forestry vehicles and their impacts on soil physical properties</article-title>. <source>Soil Till. Res.</source> <volume>79</volume>, <fpage>207</fpage>&#x02013;<lpage>219</lpage>. <pub-id pub-id-type="doi">10.1016/j.still.2004.07.009</pub-id></citation>
</ref>
<ref id="B17">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Jankovsk&#x000FD;</surname> <given-names>M.</given-names></name> <name><surname>Allman</surname> <given-names>M.</given-names></name> <name><surname>Allmanov&#x000E1;</surname> <given-names>Z.</given-names></name> <name><surname>Feren&#x0010D;&#x000ED;k</surname> <given-names>M.</given-names></name> <name><surname>Vl&#x0010D;kov&#x000E1;</surname> <given-names>M.</given-names></name></person-group> (<year>2019</year>). <article-title>Changes of key soil parameters five years after forest harvesting suggest slow regeneration of disturbed soil</article-title>. <source>J. Sustain. For.</source> <volume>38</volume>, <fpage>369</fpage>&#x02013;<lpage>380</lpage>. <pub-id pub-id-type="doi">10.1080/10549811.2018.1549500</pub-id></citation>
</ref>
<ref id="B18">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Jod&#x00142;owski</surname> <given-names>K.</given-names></name> <name><surname>Kalinowski</surname> <given-names>M.</given-names></name></person-group> (<year>2018</year>). <article-title>Current possibilities of mechanised logging in mountain areas</article-title>. <source>For. Res. Pap.</source> <volume>79</volume>, <fpage>365</fpage>&#x02013;<lpage>375</lpage>. <pub-id pub-id-type="doi">10.2478/frp-2018-0037</pub-id></citation>
</ref>
<ref id="B19">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Jourgholami</surname> <given-names>M.</given-names></name> <name><surname>Soltanpour</surname> <given-names>S.</given-names></name> <name><surname>Etehadi Abari</surname> <given-names>M.</given-names></name> <name><surname>Zenner</surname> <given-names>E. K.</given-names></name></person-group> (<year>2014</year>). <article-title>Influence of slope on physical soil disturbance due to farm tractor forwarding in a Hyrcanian forest of northern Iran</article-title>. <source>IForest Biogeosci. For.</source> <volume>7</volume>:<fpage>342</fpage>. <pub-id pub-id-type="doi">10.3832/ifor1141-007</pub-id><pub-id pub-id-type="pmid">17959540</pub-id></citation></ref>
<ref id="B20">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Labelle</surname> <given-names>E. R.</given-names></name> <name><surname>Poltorak</surname> <given-names>B. J.</given-names></name> <name><surname>Jaeger</surname> <given-names>D.</given-names></name></person-group> (<year>2019</year>). <article-title>The role of brush mats in mitigating machine-induced soil disturbances: an assessment using absolute and relative soil bulk density and penetration resistance</article-title>. <source>Can. J. For. Res.</source> <volume>49</volume>, <fpage>164</fpage>&#x02013;<lpage>178</lpage>. <pub-id pub-id-type="doi">10.1139/cjfr-2018-0324</pub-id></citation>
</ref>
<ref id="B21">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Lang</surname> <given-names>A. J.</given-names></name> <name><surname>Cristan</surname> <given-names>R.</given-names></name> <name><surname>Aust</surname> <given-names>W. M.</given-names></name> <name><surname>Bolding</surname> <given-names>M. C.</given-names></name> <name><surname>Strahm</surname> <given-names>B. D.</given-names></name> <name><surname>Vance</surname> <given-names>E. D.</given-names></name> <etal/></person-group>. (<year>2016</year>). <article-title>Long-term effects of wet and dry site harvesting on soil physical properties mitigated by mechanical site preparation in coastal plain loblolly pine (<italic>Pinus taeda</italic>) plantations</article-title>. <source>For. Ecol. Manag.</source> <volume>359</volume>, <fpage>162</fpage>&#x02013;<lpage>173</lpage>. <pub-id pub-id-type="doi">10.1016/j.foreco.2015.09.034</pub-id></citation>
</ref>
<ref id="B22">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Latterini</surname> <given-names>F.</given-names></name> <name><surname>Dyderski</surname> <given-names>M. K.</given-names></name> <name><surname>Horodecki</surname> <given-names>P.</given-names></name> <name><surname>Venanzi</surname> <given-names>R.</given-names></name> <name><surname>Picchio</surname> <given-names>R.</given-names></name> <name><surname>Jagodzi&#x00144;ski</surname> <given-names>A. M.</given-names></name></person-group> (<year>2024a</year>). <article-title>Evaluating small-scale harvesting disturbance to the forest soil in Mediterranean beech high forests</article-title>. <source>Land Degrad. Dev.</source> 1&#x02013;9. <pub-id pub-id-type="doi">10.1002/ldr.5228</pub-id></citation>
</ref>
<ref id="B23">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Latterini</surname> <given-names>F.</given-names></name> <name><surname>Horodecki</surname> <given-names>P.</given-names></name> <name><surname>Dyderski</surname> <given-names>M. K.</given-names></name> <name><surname>Scarfone</surname> <given-names>A.</given-names></name> <name><surname>Venanzi</surname> <given-names>R.</given-names></name> <name><surname>Picchio</surname> <given-names>R.</given-names></name> <etal/></person-group>. (<year>2024b</year>). <article-title>Mediterranean beech forests: thinning and ground-based skidding are found to alter microarthropod biodiversity with no effect on litter decomposition rate</article-title>. <source>For. Ecol. Manag.</source> <volume>569</volume>:<fpage>122160</fpage>. <pub-id pub-id-type="doi">10.1016/j.foreco.2024.122160</pub-id></citation>
</ref>
<ref id="B24">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Latterini</surname> <given-names>F.</given-names></name> <name><surname>Venanzi</surname> <given-names>R.</given-names></name> <name><surname>Picchio</surname> <given-names>R.</given-names></name> <name><surname>Jagodzi&#x00144;ski</surname> <given-names>A. M.</given-names></name></person-group> (<year>2023</year>). <article-title>Short-term physicochemical and biological impacts on soil after forest logging in Mediterranean broadleaf forests: 15 years of field studies summarised by a data synthesis under the meta-analytic framework</article-title>. <source>For. Int. J. For. Res.</source> <volume>96</volume>, <fpage>547</fpage>&#x02013;<lpage>560</lpage>. <pub-id pub-id-type="doi">10.1093/forestry/cpac060</pub-id></citation>
</ref>
<ref id="B25">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Macr&#x000ED;</surname> <given-names>G.</given-names></name> <name><surname>Rossi</surname> <given-names>A. D.</given-names></name> <name><surname>Papandrea</surname> <given-names>S.</given-names></name> <name><surname>Micalizzi</surname> <given-names>F.</given-names></name> <name><surname>Russo</surname> <given-names>D.</given-names></name> <name><surname>Settineri</surname> <given-names>G.</given-names></name></person-group> (<year>2017</year>). <article-title>Evaluation of soil compaction caused by passages of farm tractor in a forest in southern Italy</article-title>. <source>Agron. Res.</source> <volume>15</volume>, <fpage>478</fpage>&#x02013;<lpage>489</lpage>.</citation>
</ref>
<ref id="B26">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Malvar</surname> <given-names>M. C.</given-names></name> <name><surname>Silva</surname> <given-names>F. C.</given-names></name> <name><surname>Prats</surname> <given-names>S. A.</given-names></name> <name><surname>Vieira</surname> <given-names>D. C. S.</given-names></name> <name><surname>Coelho</surname> <given-names>C. O. A.</given-names></name> <name><surname>Keizer</surname> <given-names>J. J.</given-names></name></person-group> (<year>2017</year>). <article-title>Short-term effects of post-fire salvage logging on runoff and soil erosion</article-title>. <source>For. Ecol. Manag.</source> <volume>400</volume>, <fpage>555</fpage>&#x02013;<lpage>567</lpage>. <pub-id pub-id-type="doi">10.1016/j.foreco.2017.06.031</pub-id></citation>
</ref>
<ref id="B27">
<citation citation-type="web"><person-group person-group-type="author"><collab>Ministry of Agriculture and Rural Development of the Slovak Republic</collab></person-group> (<year>2022</year>). <source>Green Report</source>. Available at: <ext-link ext-link-type="uri" xlink:href="https://www.mpsr.sk/lesne-hospodarstvo/123">https://www.mpsr.sk/lesne-hospodarstvo/123</ext-link> (accessed January 26, 2024).</citation>
</ref>
<ref id="B28">
<citation citation-type="web"><person-group person-group-type="author"><name><surname>Mologni</surname> <given-names>O.</given-names></name> <name><surname>Dyson</surname> <given-names>P.</given-names></name> <name><surname>Amishev</surname> <given-names>D.</given-names></name> <name><surname>Proto</surname> <given-names>A. R.</given-names></name> <name><surname>Zimbalatti</surname> <given-names>G.</given-names></name> <name><surname>Cavalli</surname> <given-names>R.</given-names></name> <etal/></person-group>. (<year>2018</year>). <article-title>Tensile force monitoring on large winch-assist forwarders operating in British Columbia</article-title>. <source>Croat. J. Eng.</source> <volume>39</volume>, <fpage>193</fpage>&#x02013;<lpage>204</lpage>. Available at: <ext-link ext-link-type="uri" xlink:href="https://hrcak.srce.hr/204187">https://hrcak.srce.hr/204187</ext-link></citation>
</ref>
<ref id="B29">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Mologni</surname> <given-names>O.</given-names></name> <name><surname>Nance</surname> <given-names>E. D. T.</given-names></name> <name><surname>Lyons</surname> <given-names>C. K.</given-names></name> <name><surname>Marchi</surname> <given-names>L.</given-names></name> <name><surname>Grigolato</surname> <given-names>S.</given-names></name> <name><surname>Cavalli</surname> <given-names>R.</given-names></name> <etal/></person-group>. (<year>2021</year>). <article-title>Cable tensile forces associated to winch design in tethered harvesting operations: a case study from the Pacific North West</article-title>. <source>Forests</source> <volume>12</volume>:<fpage>827</fpage>. <pub-id pub-id-type="doi">10.3390/f12070827</pub-id></citation>
</ref>
<ref id="B30">
<citation citation-type="web"><person-group person-group-type="author"><name><surname>Najafi</surname> <given-names>A.</given-names></name> <name><surname>Solgi</surname> <given-names>A.</given-names></name></person-group> (<year>2010</year>). <article-title>Assessing site disturbance using two ground survey methods in a mountain forest</article-title>. <source>Croat. J. For. Eng.</source> <volume>31</volume>, <fpage>47</fpage>&#x02013;<lpage>55</lpage>. Available at: <ext-link ext-link-type="uri" xlink:href="https://hrcak.srce.hr/56927">https://hrcak.srce.hr/56927</ext-link></citation>
</ref>
<ref id="B31">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Picchio</surname> <given-names>R.</given-names></name> <name><surname>Mederski</surname> <given-names>P. S.</given-names></name> <name><surname>Tavankar</surname> <given-names>F.</given-names></name></person-group> (<year>2020</year>). <article-title>How and how much, do harvesting activities affect forest soil, regeneration and stands?</article-title> <source>Curr. For. Rep.</source> <volume>6</volume>, <fpage>115</fpage>&#x02013;<lpage>128</lpage>. <pub-id pub-id-type="doi">10.1007/s40725-020-00113-8</pub-id></citation>
</ref>
<ref id="B32">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Raymond</surname> <given-names>K.</given-names></name></person-group> (<year>2010</year>). <article-title>Innovative harvesting solutions: a step change harvesting research programme</article-title>. <source>N. Z. J. For.</source> <volume>55</volume>, <fpage>4</fpage>&#x02013;<lpage>9</lpage>.<pub-id pub-id-type="pmid">37842845</pub-id></citation></ref>
<ref id="B33">
<citation citation-type="book"><person-group person-group-type="author"><name><surname>Schad</surname> <given-names>P.</given-names></name></person-group> (<year>2016</year>). <article-title>&#x0201C;The International Soil Classification System WRB, Third Edition, 2014,&#x0201D;</article-title> in <source>Novel Methods for Monitoring and Managing Land and Water Resources in Siberia</source>, eds. L. Mueller, A. K. Sheudshen, and F. Eulenstein (<publisher-loc>Cham</publisher-loc>: <publisher-name>Springer International Publishing</publisher-name>), <fpage>563</fpage>&#x02013;<lpage>571</lpage>.</citation>
</ref>
<ref id="B34">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Scheer</surname> <given-names>L.</given-names></name> <name><surname>Sedm&#x000E1;k</surname> <given-names>R.</given-names></name></person-group> (<year>2010</year>). <source>Biometria (Biometry)</source>. Zvolen: Vydav. Tech. Univ. Vo Zvolene.</citation>
</ref>
<ref id="B35">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Sohrabi</surname> <given-names>H.</given-names></name> <name><surname>Jourgholami</surname> <given-names>M.</given-names></name> <name><surname>Labelle</surname> <given-names>E. R.</given-names></name></person-group> (<year>2022</year>). <article-title>The effect of forest floor on soil microbial and enzyme indices after forest harvesting operations in Hyrcanian deciduous forests</article-title>. <source>Eur. J. For. Res.</source> <volume>141</volume>, <fpage>1013</fpage>&#x02013;<lpage>1027</lpage>. <pub-id pub-id-type="doi">10.1007/s10342-022-01486-0</pub-id></citation>
</ref>
<ref id="B36">
<citation citation-type="web"><person-group person-group-type="author"><name><surname>Solgi</surname> <given-names>A.</given-names></name> <name><surname>Naghdi</surname> <given-names>R.</given-names></name> <name><surname>Tsioras</surname> <given-names>P. A.</given-names></name> <name><surname>Nikooy</surname> <given-names>M.</given-names></name></person-group> (<year>2015</year>). <article-title>Soil compaction and porosity changes caused during the operation of Timberjack 450C Skidder in Northern Iran</article-title>. <source>Croat. J. Eng.</source> <volume>36</volume>, <fpage>217</fpage>&#x02013;<lpage>225</lpage>. Available at: <ext-link ext-link-type="uri" xlink:href="https://hrcak.srce.hr/151783">https://hrcak.srce.hr/151783</ext-link></citation>
</ref>
<ref id="B37">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Sommerfeld</surname> <given-names>A.</given-names></name> <name><surname>Rammer</surname> <given-names>W.</given-names></name> <name><surname>Heurich</surname> <given-names>M.</given-names></name> <name><surname>Hilmers</surname> <given-names>T.</given-names></name> <name><surname>M&#x000FC;ller</surname> <given-names>J.</given-names></name> <name><surname>Seidl</surname> <given-names>R.</given-names></name></person-group> (<year>2021</year>). <article-title>Do bark beetle outbreaks amplify or dampen future bark beetle disturbances in Central Europe?</article-title> <source>J. Ecol.</source> <volume>109</volume>, <fpage>737</fpage>&#x02013;<lpage>749</lpage>. <pub-id pub-id-type="doi">10.1111/1365-2745.13502</pub-id><pub-id pub-id-type="pmid">33664526</pub-id></citation></ref>
<ref id="B38">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Stampfer</surname> <given-names>K.</given-names></name> <name><surname>Visser</surname> <given-names>R.</given-names></name> <name><surname>Kanzian</surname> <given-names>C.</given-names></name></person-group> (<year>2006</year>). <article-title>Cable corridor installation times for European Yarders</article-title>. <source>Int. J. For. Eng.</source> <volume>17</volume>, <fpage>71</fpage>&#x02013;<lpage>77</lpage>. <pub-id pub-id-type="doi">10.1080/14942119.2006.10702536</pub-id></citation>
</ref>
<ref id="B39">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Tavankar</surname> <given-names>F.</given-names></name> <name><surname>Bonyad</surname> <given-names>A. E.</given-names></name> <name><surname>Nikooy</surname> <given-names>M.</given-names></name> <name><surname>Picchio</surname> <given-names>R.</given-names></name> <name><surname>Venanzi</surname> <given-names>R.</given-names></name> <name><surname>Calienno</surname> <given-names>L.</given-names></name></person-group> (<year>2017</year>). <article-title>Damages to soil and tree species by cable-skidding in Caspian Forests of Iran</article-title>. <source>For. Syst.</source> <volume>26</volume>:<fpage>11</fpage>. <pub-id pub-id-type="doi">10.5424/fs/2017261-09100</pub-id></citation>
</ref>
<ref id="B40">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Tsioras</surname> <given-names>P. A.</given-names></name> <name><surname>Rottensteiner</surname> <given-names>C.</given-names></name> <name><surname>Stampfer</surname> <given-names>K.</given-names></name></person-group> (<year>2014</year>). <article-title>Wood harvesting accidents in the Austrian State Forest Enterprise 2000&#x02013;2009</article-title>. <source>Saf. Sci.</source> <volume>62</volume>, <fpage>400</fpage>&#x02013;<lpage>408</lpage>. <pub-id pub-id-type="doi">10.1016/j.ssci.2013.09.016</pub-id></citation>
</ref>
<ref id="B41">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Varol</surname> <given-names>T.</given-names></name> <name><surname>Emir</surname> <given-names>T.</given-names></name> <name><surname>Akgul</surname> <given-names>M.</given-names></name> <name><surname>Ozel</surname> <given-names>H. B.</given-names></name> <name><surname>Acar</surname> <given-names>H. H.</given-names></name> <name><surname>Cetin</surname> <given-names>M.</given-names></name></person-group> (<year>2020</year>). <article-title>Impacts of small-scale mechanized logging equipment on soil compaction in forests</article-title>. <source>J. Soil Sci. Plant Nutr.</source> <volume>20</volume>, <fpage>953</fpage>&#x02013;<lpage>963</lpage>. <pub-id pub-id-type="doi">10.1007/s42729-020-00182-5</pub-id></citation>
</ref>
<ref id="B42">
<citation citation-type="web"><person-group person-group-type="author"><name><surname>Visser</surname> <given-names>R.</given-names></name> <name><surname>Stampfer</surname> <given-names>K.</given-names></name></person-group> (<year>2015</year>). <article-title>Expanding ground-based harvesting onto steep terrain: a review</article-title>. <source>Croat. J. Eng.</source> <volume>36</volume>, <fpage>321</fpage>&#x02013;<lpage>331</lpage>. Available at: <ext-link ext-link-type="uri" xlink:href="https://hrcak.srce.hr/151831">https://hrcak.srce.hr/151831</ext-link></citation>
</ref>
<ref id="B43">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Visser</surname> <given-names>R. M.</given-names></name> <name><surname>Spinelli</surname> <given-names>R.</given-names></name></person-group> (<year>2023</year>). <article-title>Benefits and limitations of winch-assist technology for skidding operations</article-title>. <source>Forests</source> <volume>14</volume>:<fpage>296</fpage>. <pub-id pub-id-type="doi">10.3390/f14020296</pub-id></citation>
</ref>
<ref id="B44">
<citation citation-type="journal"><person-group person-group-type="author"><name><surname>Williamson</surname> <given-names>J. R.</given-names></name> <name><surname>Neilsen</surname> <given-names>W. A.</given-names></name></person-group> (<year>2000</year>). <article-title>The influence of forest site on rate and extent of soil compaction and profile disturbance of skid trails during ground-based harvesting</article-title>. <source>Can. J. For. Res.</source> <volume>30</volume>, <fpage>1196</fpage>&#x02013;<lpage>1205</lpage>. <pub-id pub-id-type="doi">10.1139/x00-041</pub-id></citation>
</ref>
</ref-list>
</back>
</article>