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<journal-meta>
<journal-id journal-id-type="publisher-id">Front. Environ. Sci.</journal-id>
<journal-title>Frontiers in Environmental Science</journal-title>
<abbrev-journal-title abbrev-type="pubmed">Front. Environ. Sci.</abbrev-journal-title>
<issn pub-type="epub">2296-665X</issn>
<publisher>
<publisher-name>Frontiers Media S.A.</publisher-name>
</publisher>
</journal-meta>
<article-meta>
<article-id pub-id-type="publisher-id">1626165</article-id>
<article-id pub-id-type="doi">10.3389/fenvs.2025.1626165</article-id>
<article-categories>
<subj-group subj-group-type="heading">
<subject>Environmental Science</subject>
<subj-group>
<subject>Systematic Review</subject>
</subj-group>
</subj-group>
</article-categories>
<title-group>
<article-title>Uneven understanding of morphological impacts of dams: upstream vs. downstream</article-title>
<alt-title alt-title-type="left-running-head">Khaleghi and Nosrati</alt-title>
<alt-title alt-title-type="right-running-head">
<ext-link ext-link-type="uri" xlink:href="https://doi.org/10.3389/fenvs.2025.1626165">10.3389/fenvs.2025.1626165</ext-link>
</alt-title>
</title-group>
<contrib-group>
<contrib contrib-type="author" corresp="yes">
<name>
<surname>Khaleghi</surname>
<given-names>Somaiyeh</given-names>
</name>
<xref ref-type="corresp" rid="c001">&#x2a;</xref>
<uri xlink:href="https://loop.frontiersin.org/people/3056155/overview"/>
<role content-type="https://credit.niso.org/contributor-roles/supervision/"/>
<role content-type="https://credit.niso.org/contributor-roles/formal-analysis/"/>
<role content-type="https://credit.niso.org/contributor-roles/writing-original-draft/"/>
<role content-type="https://credit.niso.org/contributor-roles/Writing - review &#x26; editing/"/>
<role content-type="https://credit.niso.org/contributor-roles/software/"/>
<role content-type="https://credit.niso.org/contributor-roles/data-curation/"/>
<role content-type="https://credit.niso.org/contributor-roles/investigation/"/>
<role content-type="https://credit.niso.org/contributor-roles/methodology/"/>
<role content-type="https://credit.niso.org/contributor-roles/project-administration/"/>
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</contrib>
<contrib contrib-type="author">
<name>
<surname>Nosrati</surname>
<given-names>Kazem</given-names>
</name>
<role content-type="https://credit.niso.org/contributor-roles/conceptualization/"/>
<role content-type="https://credit.niso.org/contributor-roles/Writing - review &#x26; editing/"/>
<role content-type="https://credit.niso.org/contributor-roles/methodology/"/>
</contrib>
</contrib-group>
<aff>
<institution>Department of Physical Geography, Faculty of Earth Sciences, Shahid Beheshti University</institution>, <addr-line>Tehran</addr-line>, <country>Iran</country>
</aff>
<author-notes>
<fn fn-type="edited-by">
<p>
<bold>Edited by:</bold> <ext-link ext-link-type="uri" xlink:href="https://loop.frontiersin.org/people/250016/overview">YoonKyung Cha</ext-link>, University of Seoul, Republic of Korea</p>
</fn>
<fn fn-type="edited-by">
<p>
<bold>Reviewed by:</bold> <ext-link ext-link-type="uri" xlink:href="https://loop.frontiersin.org/people/2184002/overview">Michael Pollock</ext-link>, NOAA Fisheries, United States</p>
<p>
<ext-link ext-link-type="uri" xlink:href="https://loop.frontiersin.org/people/2421581/overview">Chrysoula Ntislidou</ext-link>, Aristotle University of Thessaloniki, Greece</p>
</fn>
<corresp id="c001">&#x2a;Correspondence: Somaiyeh Khaleghi, <email>s_khaleghi@sbu.ac.ir</email>
</corresp>
</author-notes>
<pub-date pub-type="epub">
<day>21</day>
<month>10</month>
<year>2025</year>
</pub-date>
<pub-date pub-type="collection">
<year>2025</year>
</pub-date>
<volume>13</volume>
<elocation-id>1626165</elocation-id>
<history>
<date date-type="received">
<day>10</day>
<month>05</month>
<year>2025</year>
</date>
<date date-type="accepted">
<day>13</day>
<month>10</month>
<year>2025</year>
</date>
</history>
<permissions>
<copyright-statement>Copyright &#xa9; 2025 Khaleghi and Nosrati.</copyright-statement>
<copyright-year>2025</copyright-year>
<copyright-holder>Khaleghi and Nosrati</copyright-holder>
<license xlink:href="http://creativecommons.org/licenses/by/4.0/">
<p>This is an open-access article distributed under the terms of the Creative Commons Attribution License (CC BY). The use, distribution or reproduction in other forums is permitted, provided the original author(s) and the copyright owner(s) are credited and that the original publication in this journal is cited, in accordance with accepted academic practice. No use, distribution or reproduction is permitted which does not comply with these terms.</p>
</license>
</permissions>
<abstract>
<p>Despite the extensive knowledge on dam-induced river adjustments, systematic comparisons and integrated syntheses of both upstream and downstream morphological effects are still lacking. In this study, we systematically reviewed all available studies on these adjustments from the SCOPUS and WOS databases. We found that the majority of the 95 analyzed papers (80%) focused exclusively on downstream river channel changes, such as channel incision and narrowing. In contrast, upstream adjustments, including channel widening and aggradation, were addressed in only 16% of the articles, and their associated ecological impacts remain largely understudied. Most of the reviewed studies used remote sensing methods to analyze river morphological adjustments in temperate climates and single-thread rivers. This review highlights the limited understanding of river morphological changes occurring upstream from dams and their ecological consequences. Addressing this gap in further research is crucial for informing future river management practices aimed at mitigating the impacts of dams.</p>
</abstract>
<kwd-group>
<kwd>systematic review</kwd>
<kwd>dam</kwd>
<kwd>channel morphology</kwd>
<kwd>upstream</kwd>
<kwd>downstream</kwd>
</kwd-group>
<counts>
<page-count count="11"/>
</counts>
<custom-meta-wrap>
<custom-meta>
<meta-name>section-at-acceptance</meta-name>
<meta-value>Freshwater Science</meta-value>
</custom-meta>
</custom-meta-wrap>
</article-meta>
</front>
<body>
<sec sec-type="intro" id="s1">
<title>1 Introduction</title>
<p>Dams are one of the oldest man-made infrastructures constructed for flood control, irrigation, water supply and navigation (<xref ref-type="bibr" rid="B66">Zhang and Gu, 2023</xref>). These structures can profoundly alter both hydrological and sediment load regimes of rivers (e.g., <xref ref-type="bibr" rid="B58">Williams et al., 1984</xref>; <xref ref-type="bibr" rid="B50">Poff et al., 1997</xref>; <xref ref-type="bibr" rid="B21">Knighton, 1998</xref>; <xref ref-type="bibr" rid="B4">Brandt, 2000</xref>; <xref ref-type="bibr" rid="B14">Grant et al., 2003</xref>; <xref ref-type="bibr" rid="B43">Nilsson et al., 2005</xref>; <xref ref-type="bibr" rid="B12">Graf, 2006</xref>; <xref ref-type="bibr" rid="B13">Grant, 2012</xref>). While dams have existed for thousands of years, the majority have been constructed in the last 60&#xa0;years, resulting in around 58,000 large dams currently operating worldwide (<xref ref-type="bibr" rid="B41">Mulligan et al., 2020</xref>). The International Commission on Large Dams (ICOLD) defined large dam as &#x201c;a dam with 15&#xa0;m or greater in height (measured from the lowest point of foundation to top of dam) or a dam between 5 and 15&#xa0;m impounding more than 3 million cubic metre&#x201d; (<xref ref-type="bibr" rid="B66">Zhang and Gu, 2023</xref>).</p>
<p>Dams are widely documented as factors triggering diverse morphological adjustments of river channels and floodplains (e.g., <xref ref-type="bibr" rid="B45">Petts, 1979</xref>; <xref ref-type="bibr" rid="B47">1980b</xref>; <xref ref-type="bibr" rid="B21">Knighton, 1998</xref>; <xref ref-type="bibr" rid="B48">Petts and Gurnell, 2005</xref>; <xref ref-type="bibr" rid="B10">Gordon and Meentemeyer, 2006</xref>; <xref ref-type="bibr" rid="B54">Takahashi and Nakamura, 2011</xref>; <xref ref-type="bibr" rid="B13">Grant, 2012</xref>; <xref ref-type="bibr" rid="B36">Ma et al., 2012</xref>; <xref ref-type="bibr" rid="B5">Chong et al., 2021</xref>), with significant implications for ecological functioning of rivers (<xref ref-type="bibr" rid="B26">Ligon et al., 1995</xref>), flood risk in river valleys (<xref ref-type="bibr" rid="B31">Liro et al., 2020</xref>; <xref ref-type="bibr" rid="B37">Ma et al., 2022</xref>), and bottomland management (<xref ref-type="bibr" rid="B30">Liro, 2019</xref>).</p>
<p>These morphological changes are generally categorized into those occurring upstream and downstream of dams (<xref ref-type="bibr" rid="B27">Liro, 2014</xref>). The downstream morphological effects of dams (e.g., <xref ref-type="bibr" rid="B4">Brandt, 2000</xref>; <xref ref-type="bibr" rid="B48">Petts and Gurnell, 2005</xref>; <xref ref-type="bibr" rid="B13">Grant, 2012</xref>) are substantially better understood than their upstream counterparts (<xref ref-type="bibr" rid="B27">Liro, 2014</xref>; <xref ref-type="bibr" rid="B30">Liro, 2019</xref>; <xref ref-type="bibr" rid="B31">Liro et al., 2020</xref>). Despite the abundance of research on dam-induced morphological adjustments, a global synthesis comparing upstream and downstream impacts remains lacking. Such a synthesis is critical for building a comprehensive understanding of the historical, current, and projected future consequences of dams on riverine ecosystems. Furthermore, it may guide the development of targeted management strategies for dam-affected river segments, especially those located in regions with additional natural or anthropogenic pressures (e.g., regulated rivers in arid or semi-arid zones). In such systems, dam-induced morphological alterations&#x2014;such as channel incision&#x2014;can exacerbate vulnerability to future climate change impacts.</p>
<p>In this study, we conducted a systematic review of existing literature addressing the upstream and downstream morphological effects of dams on rivers. The provided synthesis can help better understand the patterns of dam-induced channel adjustment in various rivers and support future river management to reduce ecological and economic losses resulting from them. Based on the analysis of 95 papers, we: (i) synthesized the dominant patterns of dam-induced morphological changes across global rivers with varying climatic and morphological characteristics, and (ii) identified main knowledge gaps in our recognition of the above problem, suggesting future research directions allowing for narrowing them.</p>
</sec>
<sec sec-type="materials|methods" id="s2">
<title>2 Materials and methods</title>
<p>This study was conducted using the principles of the Preferred Reporting Items for Systematic Reviews and Meta-Analyses (PRISMA) (<xref ref-type="bibr" rid="B44">Page et al., 2021</xref>) and analyzed all available English-written papers related to the spatial and temporal patterns of dam-induced changes in river channel morphology worldwide.</p>
<sec id="s2-1">
<title>2.1 Information sources</title>
<p>The review was performed utilizing the SCOPUS and Web of Science (WOS) databases to assess relevant studies published between 1974 and 2023.</p>
</sec>
<sec id="s2-2">
<title>2.2 Search strategy</title>
<p>The initial search was conducted using the Boolean operators &#x201c;AND&#x201d; and &#x201c;OR,&#x201d; combined with keywords related to the subject area, such as &#x201c;channel morphology (river morphology, channel cross-section, channel erosion, channel aggradation and riverbank erosion, channel degradation, incision, narrowing, widening, bed level),&#x201d; &#x201c;dam (reservoir, hydroelectric power plant),&#x201d; and &#x201c;hydrology (stream flow, peak flow, discharge, sediment yield, sediment load)&#x201d; etc. to identify and select papers for review.</p>
</sec>
<sec id="s2-3">
<title>2.3 Eligibility criteria</title>
<p>Eligibility of references was determined based on predefined inclusion and exclusion criteria. Only original research articles published in English in peer-reviewed journals focusing on river morphological responses to dam construction were included. Books, conference papers, review articles, and presentations about channel changes were excluded from the study. In the initial research, after removing duplicate papers using EndNote X6, based on the title assessment, some papers were excluded. Subsequently, papers with unrelated topic, duplicate content, and manuscripts with lacking clear methodology, evaluation criteria, or results were also excluded. Importantly, this study focuses exclusively on the effects of dams on river morphology; other anthropogenic influences such as land use changes, mining, or urbanization were not considered.</p>
</sec>
<sec id="s2-4">
<title>2.4 Selection process</title>
<p>The preliminary search results were imported into Endnote X6, and duplicates were removed. Two independent authors then screened the titles and abstracts of all the papers and omitted some irrelevant papers based on the eligibility criteria. Afterward, the full texts of the remaining papers were downloaded and thoroughly reviewed to ensure that they meet the criteria for selection. Finally, eligible papers were systematically analyzed to extract relevant data. The following information was collected from each article: fist author, year of publication, river name, geographical location, timeframe of observed changes, dam name, location relative to dam (upstream or downstream), climatic zone, river pattern, data collection method, and observed morphological changes.</p>
<p>Finally, the papers were classified according to type of dam-induced morphological change (e.g., widening, narrowing, deepening, shallowing), position relative to the dam (upstream or downstream), river pattern (single-thread, multi-thread), location of the study area (continent, climatic zone [K&#xf6;ppen-Geiger (<ext-link ext-link-type="uri" xlink:href="https://koeppen-geiger.vu-wien.ac.at/">https://koeppen-geiger.vu-wien.ac.at/</ext-link>)] and method used for quantifying morphological changes (e.g., topographic measurements, remote sensing material, hydrological data) (see <xref ref-type="sec" rid="s12">Supplementary Table S1</xref>).</p>
</sec>
</sec>
<sec sec-type="results" id="s3">
<title>3 Results</title>
<sec id="s3-1">
<title>3.1 Study selection</title>
<p>A combined search of titles, abstracts, and keywords was conducted. First, the literature search was conducted using the SCOPUS database, which resulted in 172 articles found. Then, an additional search was carried out using the Web of Science database, resulting in 118 articles indicated. In total, 290 unique records were identified across both databases (n &#x3d; 290). Subsequently, the database was reviewed to remove duplicates (n &#x3d; 47) and articles not directly related to the study topic (n &#x3d; 62). During the subsequent phase, a total of 181 records were assessed based on the title and abstract content, taking into account the exclusion criteria such as irrelevant keywords (e.g., check dam, other human intervention, experimental and flume studies). As a result, 59 articles were eliminated. The whole texts of the remaining records (n &#x3d; 122) were evaluated to determine their eligibility. From 122 records, the articles which did not have enough data on river morphological adjustments (n &#x3d; 21) and those have the effect of dam combined with other human interventions (n &#x3d; 6) were excluded (<italic>n</italic> &#x3d; 27). Finally, 95 articles met all inclusion criteria and were included in the systematic review (<xref ref-type="fig" rid="F1">Figure 1</xref>).</p>
<fig id="F1" position="float">
<label>FIGURE 1</label>
<caption>
<p>PRISMA workflow applied in this work to literature extraction.</p>
</caption>
<graphic xlink:href="fenvs-13-1626165-g001.tif">
<alt-text content-type="machine-generated">Flowchart depicting the process of study identification and inclusion via databases. It includes three stages: Identification, Screening, and Included. Initially, 290 peer-reviewed articles from Scopus and WOS are identified. Records are reduced after removing duplicates and other reasons, resulting in 181 screened records. After excluding 59 for unrelated keywords, 122 reports are sought, with no retrieval discrepancies. Eligibility assessment leads to exclusion of 27 reports, due to insufficient data and combined effects, resulting in 95 studies included in the review.</alt-text>
</graphic>
</fig>
</sec>
<sec id="s3-2">
<title>3.2 Publication trend and study location</title>
<p>An analysis of the existing literature revealed a notable increase in the number of publications addressing dam-induced morphological changes between 1974 and 2023 (<xref ref-type="fig" rid="F2">Figure 2</xref>). This is parallel to the increase in dam construction, so that according to <xref ref-type="bibr" rid="B66">Zhang and Gu (2023)</xref>, over the past three&#xa0;decades, dams have increased especially in developing countries in Asia and South America. The geographic distribution of the 95 reviewed studies is presented in <xref ref-type="fig" rid="F3">Figure 3</xref>. Among these, 57 studies (60%) were conducted in Asia, followed by 16 (17%) in Europe and North America, 4 (4%) in South America, and 2 (2%) in Africa (<xref ref-type="table" rid="T1">Table 1</xref>). Also, number of the rivers and dams located in countries is shown in <xref ref-type="table" rid="T1">Table 1</xref>. The most frequently cases are in Asia from China (38 papers), primarily related to the Yangtze and Yellow Rivers (<xref ref-type="table" rid="T1">Table 1</xref>; <xref ref-type="sec" rid="s12">Supplementary Table S1</xref>). The Yangtze River basin has area of 1.8 &#xd7; 10<sup>6</sup>&#xa0;km<sup>2</sup> with 6,300&#xa0;km river length and the Yellow River basin is 752,443&#xa0;km<sup>2</sup> with 3,471&#xa0;km length (<xref ref-type="sec" rid="s12">Supplementary Table S2</xref>). These rivers are among longest river in the world which have several dams built on them and their tributaries (<xref ref-type="sec" rid="s12">Supplementary Table S1</xref>). Regarding climatic distribution, 59 studies (62%) were conducted in temperate climates, followed by 21 (22%) in semi-arid, 11 (12%) in continental, and 4 (4%) in tropical climates (<xref ref-type="fig" rid="F4">Figure 4</xref>; <xref ref-type="sec" rid="s12">Supplementary Table S1</xref>). Therefore, these studies are scattered across different climatic regions. The K&#xf6;ppen climate classification scheme divides climates into five main climate groups: A (tropical), B (arid), C (temperate), D (continental), and E (polar). Each group and subgroup is represented by a letter. The second letter indicates the seasonal precipitation type, while the third letter indicates the level of heat. According to <xref ref-type="sec" rid="s12">Supplementary Table S1</xref>, the climate of study locations define as: Aw: Tropical wet and dry (savanna climate); BSh: Hot semi-arid climate; BSk: Cold semi-arid climate; Cfa: Humid subtropical climate; Cfb: Temperate oceanic climate or subtropical highland climate; Csa: Hot-summer Mediterranean climate; Csb: Warm-summer Mediterranean climate; Cwa: Monsoon; Dfa: Hot-summer humid continental climate; Dfb: Warm-summer humid continental climate; Dwb: Monsoon-influenced warm-summer humid continental climate.</p>
<fig id="F2" position="float">
<label>FIGURE 2</label>
<caption>
<p>Publication time of articles used in the systematic review.</p>
</caption>
<graphic xlink:href="fenvs-13-1626165-g002.tif">
<alt-text content-type="machine-generated">Bar chart showing the number of studies from 1974 to 2023. Notable increases occur around 1993, 2005, and 2010 onwards, peaking in 2022 with twelve studies.</alt-text>
</graphic>
</fig>
<fig id="F3" position="float">
<label>FIGURE 3</label>
<caption>
<p>The location of 95 case studies used in systematic review.</p>
</caption>
<graphic xlink:href="fenvs-13-1626165-g003.tif">
<alt-text content-type="machine-generated">World map displaying various continents, including North America, South America, Europe, Africa, Asia, Australia, and Oceania. Several magenta dots are spread across these continents, indicating specific locations.</alt-text>
</graphic>
</fig>
<table-wrap id="T1" position="float">
<label>TABLE 1</label>
<caption>
<p>The reviewed case studies included reports on the rivers and dam&#x2019;s location, number, and initial channel morphology.</p>
</caption>
<table>
<thead valign="top">
<tr>
<th align="center">Continent</th>
<th align="center">Location of rivers</th>
<th align="center">Number of rivers</th>
<th align="center">Number of dams</th>
<th align="center">Upstream (U)/downstream (D) of dam</th>
<th align="center">Channel morphology<xref ref-type="table-fn" rid="Tfn1">
<sup>a</sup>
</xref>
</th>
<th align="center">Number and proportion (%) of studies</th>
</tr>
</thead>
<tbody valign="top">
<tr>
<td align="center">Asia</td>
<td align="center">China, India, Iran, Korea, Turkey, Japan, Russia, Iraq</td>
<td align="center">21</td>
<td align="center">18</td>
<td align="center">U (n &#x3d; 6), D (n &#x3d; 47), U&#x26;D (n &#x3d; 4)</td>
<td align="center">S (n &#x3d; 39), M (n &#x3d; 6), S&#x26;M (n &#x3d; 12)</td>
<td align="center">57 (60%)</td>
</tr>
<tr>
<td align="center">Europe</td>
<td align="center">Poland, Italy, Spain, England, Hungary</td>
<td align="center">11</td>
<td align="center">11</td>
<td align="center">U (n &#x3d; 5), D (n &#x3d; 10), U&#x26;D (n &#x3d; 1)</td>
<td align="center">S (n &#x3d; 14), M (n &#x3d; 2), S&#x26;M (n &#x3d; 0)</td>
<td align="center">16 (17%)</td>
</tr>
<tr>
<td align="center">North America</td>
<td align="center">USA, Mexico, Canada</td>
<td align="center">10</td>
<td align="center">10</td>
<td align="center">U (n &#x3d; 2), D (n &#x3d; 12), U&#x26;D (n &#x3d; 1)</td>
<td align="center">S (n &#x3d; 14), M (n &#x3d; 2), S&#x26;M (n &#x3d; 0)</td>
<td align="center">16 (17%)</td>
</tr>
<tr>
<td align="center">South America</td>
<td align="center">Ecuador, Brazil, Colombia, Chile</td>
<td align="center">5</td>
<td align="center">5</td>
<td align="center">U (n &#x3d; 0), D (n &#x3d; 3), U&#x26;D (n &#x3d; 1)</td>
<td align="center">S (n &#x3d; 2), M (n &#x3d; 0), S&#x26;M (n &#x3d; 2)</td>
<td align="center">4 (4%)</td>
</tr>
<tr>
<td align="center">Africa</td>
<td align="center">Mozambique, Ethiopia</td>
<td align="center">2</td>
<td align="center">2</td>
<td align="center">U (n &#x3d; 0), D (n &#x3d; 2), U&#x26;D (n &#x3d; 0)</td>
<td align="center">S (n &#x3d; 1), M (n &#x3d; 1), S&#x26;M (n &#x3d; 0)</td>
<td align="center">2 (2%)</td>
</tr>
<tr>
<td align="center">Sum</td>
<td align="center">22</td>
<td align="center">49</td>
<td align="center">46</td>
<td align="center">U (n &#x3d; 13), D (n &#x3d; 76), U&#x26;D (n &#x3d; 6)</td>
<td align="center">S (n &#x3d; 71), M (n &#x3d; 11), S&#x26;M (n &#x3d; 13)</td>
<td align="center">95 (100%)</td>
</tr>
</tbody>
</table>
<table-wrap-foot>
<fn id="Tfn1">
<label>
<sup>a</sup>
</label>
<p>S: Single-thread, M: Multi-thread, S&#x26;M: Single and Multi-thread.</p>
</fn>
</table-wrap-foot>
</table-wrap>
<fig id="F4" position="float">
<label>FIGURE 4</label>
<caption>
<p>Number and proportion of reviewed publications based on the climate of study location.</p>
</caption>
<graphic xlink:href="fenvs-13-1626165-g004.tif">
<alt-text content-type="machine-generated">Bar chart depicting the number and proportion of studies conducted in various climates. Temperate climates have the highest with 59 studies, accounting for 62%. Arid climates follow with 21 studies at 22%. Continental climates have 11 studies, representing 11%, while tropical climates have the fewest with 4 studies, comprising 4%.</alt-text>
</graphic>
</fig>
</sec>
<sec id="s3-3">
<title>3.3 Dam-induced morphological changes</title>
<p>A significant proportion of the reviewed studies (76 papers, or 80%) focused on the downstream effects of dams on river morphology, while a significantly smaller number of papers investigated the upstream effects (13 papers, or 14%) or both upstream and downstream effects (6 papers, or 6%) (<xref ref-type="table" rid="T1">Table 1</xref>). In terms of river types, most studies (71; 75%) were conducted on single-thread channels, while 11 studies (12%) focused on multi-thread systems, and 13 studies (13%) included both types. Most of the studies have been carried out in reaches located in alluvial (76.84%), alluvial and mountainous (10.53%), mountainous (7.37%), alluvial and coastal (3.16%), and coastal plains (2.11%), respectively. The shortest reach studied was 1.5&#xa0;km upstream of a dam in Smolnik, Poland, while the longest extended 1,300&#xa0;km downstream along the Lower Yangtze River, China (<xref ref-type="sec" rid="s12">Supplementary Table S1</xref>).</p>
<p>In terms of morphological adjustments, channel incision (48; 56%), narrowing (24; 28%), have been most frequently reported in the river reaches downstream from dams. Narrowing ranged between 10% and 75%. The highest amount of channel narrowing (75%) have been reported by <xref ref-type="bibr" rid="B7">del Tanago et al. (2015)</xref> in downstream of Guadalete river in Spain (<xref ref-type="sec" rid="s12">Supplementary Table S1</xref>). This study reach is single-thread in morphology and located at temperate climate (Csa). Also, <xref ref-type="bibr" rid="B59">Wu et al. (2023)</xref> stated that channel narrowing (60%) has been occurred in downstream reach of Lower Yellow River as a multi-thread river in temperate climate (Cwa). About bed-level changes, the highest incision over 5&#xa0;m reported in downstream of the single-thread and multi-thread rivers by <xref ref-type="bibr" rid="B19">Kale and Ataol (2021)</xref> in Ye&#x15f;il&#x131;rmak (Turkey) and <xref ref-type="bibr" rid="B22">Kondolf and Swanson (1993)</xref> in Stony Creek (California). In contrast with downstream, channel widening (6; 32%) and aggradation (9; 47%) have been most frequent reported in the river reaches upstream from dams (<xref ref-type="fig" rid="F5">Figures 5</xref>, <xref ref-type="fig" rid="F7">7</xref>). The highest channel widening up to 600% has occurred in upstream of Magdalena River in Colombia with tropical climate (<xref ref-type="bibr" rid="B2">Alvarado et al., 2023</xref>). Only just few papers about channel aggradation were stated but these papers didn&#x2019;t report amount of aggradation.</p>
<fig id="F5" position="float">
<label>FIGURE 5</label>
<caption>
<p>Types of dam-induced morphological changes reported in the reviewed case studies.</p>
</caption>
<graphic xlink:href="fenvs-13-1626165-g005.tif">
<alt-text content-type="machine-generated">Bar chart showing the number and percentage of studies on downstream and upstream dam effects. Categories include Incision, Aggradation, Narrowing, and Widening. Downstream dams have higher percentages for Incision (56%) and Narrowing (28%), while upstream dams have higher percentages for Aggradation (64%) and Widening (43%).</alt-text>
</graphic>
</fig>
<p>In the case of initially multi-thread rivers located upstream of dams, long-term channel narrowing and pattern simplification have been observed (e.g., <xref ref-type="bibr" rid="B61">Xu, 1990</xref>; <xref ref-type="bibr" rid="B62">Xu, 2001</xref>; <xref ref-type="bibr" rid="B63">Xu and Shi, 1997</xref>), although some channel width fluctuations may occur during the initial phase of adjustment (<xref ref-type="bibr" rid="B29">Liro, 2016</xref>). For example, <xref ref-type="bibr" rid="B28">Liro (2015)</xref>, <xref ref-type="bibr" rid="B29">Liro (2016)</xref> reconstructed that in a gravel-bed river, a rise in the downstream base level and the onset (&#x223c;first 20 years) of backwater fluctuations in upstream channel sections led to channel widening, driven by accelerated in-channel deposition, bar development, and associated bank erosion.</p>
<p>From river pattern, in the case of downstream-dam reaches a few papers reported a decrease in sinuosity (6 papers) (e.g., <xref ref-type="bibr" rid="B61">Xu, 1990</xref>; <xref ref-type="bibr" rid="B40">Morais et al., 2016</xref>; <xref ref-type="bibr" rid="B65">Yousefi, et al., 2016</xref>; <xref ref-type="bibr" rid="B19">Kale and Ataol, 2021</xref>; <xref ref-type="bibr" rid="B2">Alvarado, et al., 2023</xref>; <xref ref-type="bibr" rid="B6">Clavijo-Rivera, et al., 2023</xref>) and braiding index (4 papers) (e.g., Surian, 1999; Liro, 2017; <xref ref-type="bibr" rid="B25">Kumar, et al., 2019</xref>; <xref ref-type="bibr" rid="B23">Kong, et al., 2020</xref>), but only one study reported these effects in upstream-dam sections (<xref ref-type="sec" rid="s12">Supplementary Table S1</xref>). Eight papers noted a decrease in bar areas downstream of dams in just a few case studies, including transitions from wandering to single-thread (1 paper), braided to single-thread (1 paper), and braided to wandering (1 paper). For instance, <xref ref-type="bibr" rid="B25">Kumar et al. (2019)</xref> found that the Wazirabad Dam in India caused a decline in bar area, a reduced braiding index, and a narrower channel belt downstream of the Yamuna River.</p>
<p>After operation of dam, reducing peak discharges and sediment may lead to changes in lateral channel migration in downstream of dams (e.g., <xref ref-type="bibr" rid="B49">Phillips et al., 2005</xref>; <xref ref-type="bibr" rid="B10">Gordon and Meentemeyer, 2006</xref>; <xref ref-type="bibr" rid="B42">Musselman, 2011</xref>; <xref ref-type="bibr" rid="B51">Smith and Mohrig, 2017</xref>; <xref ref-type="bibr" rid="B23">Kong et al., 2020</xref>; <xref ref-type="bibr" rid="B32">Liu et al., 2021</xref>; <xref ref-type="bibr" rid="B17">He et al., 2022</xref>; <xref ref-type="bibr" rid="B11">Goudie, 2022</xref>). Also coarsening of channel sediment in downstream of dam was reported in few papers (e.g., <xref ref-type="bibr" rid="B49">Phillips et al., 2005</xref>; <xref ref-type="bibr" rid="B51">Smith and Mohrig, 2017</xref>; <xref ref-type="bibr" rid="B8">Gao et al., 2021</xref>; <xref ref-type="bibr" rid="B64">Yang et al., 2023</xref>). In this regard, only one paper by <xref ref-type="bibr" rid="B38">Maselli et al. (2016)</xref> stated that in Isola Serafini dam in Italy, lateral migration rate of the channel is up to 45&#xa0;m/yr upstream of the influence of backwater flow and ca.10&#xa0;m/yr at the transition from normal to backwater flow conditions (30&#xa0;km from the dam) and deposition of coarse-grained sediment.</p>
<p>It should be noted that by alteration of hydrology, sediment and morphology, vegetation cover changes. In some cases, sedimentation and encroachment of vegetation in single-thread reaches have been reported as a result of reduced flow downstream of dams (e.g., Choi et al., 2005; Asaeda and Rashid, 2012; <xref ref-type="bibr" rid="B7">del Tanago et al., 2015</xref>; <xref ref-type="bibr" rid="B64">Yang et al., 2023</xref>). Only Liro et al. (2022) demonstrated that backwater fluctuations of Ro&#x2d9;zn&#xf3;w dam on upstream of Smolnik River substantially disturb the hydrodynamics, which is lead to alterations in sedimentology, morphology, and riparian vegetation pattern. <xref ref-type="bibr" rid="B30">Liro (2019)</xref> suggested that changes in the river hydrodynamics and the deposition of fine sediments in upstream of dam, may be the factors triggering vegetation expansion. However, the same factors may eliminate plants that are not resistant to prolonged inundation and high sedimentation rate. Up to now there have been no works presenting a spatial&#x2013;temporal reconstruction of this process and its relation to the geomorphic characteristics of a given site in the backwater.</p>
<p>These morphological adjustments were most commonly reconstructed using remote sensing materials, including satellite images, aerial photos, and maps (37 papers; 39%), data from field measurements (19; 20%), and historical hydrological data (8; 8%). A high number of works (31; 33%) utilized more than one data collection method (<xref ref-type="fig" rid="F6">Figure 6</xref>). The use of remote sensing techniques has increased significantly in studies from 1992 to 2023. According to <xref ref-type="bibr" rid="B5">Chong et al. (2021)</xref>, these remote techniques are increasingly replacing field-based approaches to studying geomorphic processes and changes. After gathering data on channel morphology pre and post-dam periods, the researchers have used the statistical analysis and/or hydrological modeling for distingutiong channel changes due to dam construction.</p>
<fig id="F6" position="float">
<label>FIGURE 6</label>
<caption>
<p>Methods used for collecting data on dam-induced morphological adjustments in the reviewed case studies.</p>
</caption>
<graphic xlink:href="fenvs-13-1626165-g006.tif">
<alt-text content-type="machine-generated">Bar chart displaying the number and proportion of studies using various methods. Remote Sensing Materials leads with 37 studies (39%), followed by Few Methods at 31 (33%), Field Surveys at 19 (20%), and Analysis of Hydrological Data at 8 (8%).</alt-text>
</graphic>
</fig>
<p>Finally, it is possible that channel changes have environmental and ecosystem effects, but such effects have not been evaluated broadly in the existing studies. However, few of the reviewed papers mentioned ecological and management consequences of channel adjustments in downstream of dams and only one paper mentioned these in upstream. The work of <xref ref-type="bibr" rid="B22">Kondolf and Swanson (1993)</xref> in Stony Creek, showed that the change from a braided channel pattern to a more sinuous and meandering channel, probably reflecting an adjustment to abruptly reduced rates of bedload sediment supply from the watershed, to the decrease in flood peaks and sediment transport capacity, and possibly to prolonged bankfull flows. <xref ref-type="bibr" rid="B49">Phillips et al. (2005)</xref> has reported intensive bank erosion downstream of dam in Lower Trinity. Sanchis Ibor1 et al. (2018) reported that transformation from wandering to single thread channel pattern in Serpis river was followed by a slow vegetation encroachment, and culminated with the stabilization of channel migration. The role of vegetation has been critical in controlling floods&#x27; effectiveness, reducing river mobility and shifting, and consolidating a channel planform model adapted to the post-dam flow conditions. Wu et al. (2018) demonstrated that after construction of Zhikong Reservoir on Lhasa River, the number and area of central bars in the braided reach closest to Zhikong Dam (RS1) were increased because of main channel incision and water level drop. <xref ref-type="bibr" rid="B19">Kale and Ataol (2021)</xref> mentioned that river degradation due to the effects of dams on Ye&#x15f;il&#x131;rmak River has a major environmental problem at the historical Car&#x15f;amba Bridge and also along the riverbed of the Ye&#x15f;il&#x131;rmak River after dam construction. Agarwal et al. (2022) said that the ability of the Srinagar dam to adjust the sediment load of the event in its upstream is an immediate short-term but it creates a geomorphic disconnect in the channel between upstream and downstream reaches of the reservoir. This disconnect may have a negative effect on sediment&#x2013;water routing of the river and sediment storage. About upstream, Liro et al., (2022) mentioned that the fluctuation effects of reservoir backwater have caused the change of channel pattern from the multi-thread to the single-thread. This increased the riparian forest expansion in the stream valley and decreased the floodplain utilization potential and caused the development of homogenous and simplified morphological and vegetation patterns.</p>
</sec>
</sec>
<sec sec-type="discussion" id="s4">
<title>4 Discussion</title>
<p>A systematic review of published literature reveals that a total of 95 papers have investigated the effects of dams on river channel adjustments globally. However, this number is relatively low when compared to the vast scale of dam construction around the world. This finding is consistent with <xref ref-type="bibr" rid="B5">Chong et al. (2021)</xref>, who noted that studies addressing geomorphological changes in tropical rivers are scarce, particularly in comparison to research on flow regimes and sediment transport.</p>
<p>The present review shows that most of the studies focused on single-thread rivers and downstream reaches of dams. In terms of location and climate, major studies have been conducted in the Asian rivers and temperate climates. According to <xref ref-type="bibr" rid="B66">Zhang and Gu (2023)</xref> Asia hosts the largest number of dams globally (9,526&#xa0;dams, or 27% of the total). North and South America follow, with 23% and 21% of global dam counts, respectively. Among the Asian case studies, the Yangtze and Yellow Rivers were the most frequently studied, which is unsurprising given the Yangtze&#x2019;s status as the third-longest river in the world (6,300&#xa0;km; He et al., 2021) and the location of the Three Gorges Dam (TGD) &#x2014; the largest dam in the world (<xref ref-type="bibr" rid="B67">Zhang et al., 2016</xref>). Despite the extensive studies on dam-induced changes along the Yangtze, major rivers such as the Nile and Amazon etc. remain understudied in this context. Additionally, no case studies were found documenting dam impacts on river morphology in the Oceania continent, a finding aligned with <xref ref-type="bibr" rid="B66">Zhang and Gu (2023)</xref>, who reported that Oceania has the lowest number of dams globally.</p>
<p>Mainly, large dams have decreased the sediment transporting flows by more than 80%, which has resulted in a much lower capacity to change the channel (<xref ref-type="bibr" rid="B7">del Tanago et al., 2015</xref>). According to literature review, narrowing and incision are the dominant process in downstream of dams (<xref ref-type="fig" rid="F7">Figure 7</xref>) (e.g., Surian, 1999; Choi et al., 2005; Jian-guo et al., 2012; <xref ref-type="bibr" rid="B36">Ma et al., 2012</xref>; <xref ref-type="bibr" rid="B3">Ashouri et al., 2015</xref>; <xref ref-type="bibr" rid="B1">Adib et al., 2016</xref>; <xref ref-type="bibr" rid="B9">Gierszewski et al., 2020</xref>; <xref ref-type="bibr" rid="B68">Zhou et al., 2021</xref>; <xref ref-type="bibr" rid="B16">Han et al., 2018</xref>; <xref ref-type="bibr" rid="B56">Wang et al., 2022</xref>; <xref ref-type="bibr" rid="B59">Wu et al., 2023</xref>). Dams significantly alter river morphology by interrupting sediment transport processes and modifying hydrological patterns, which subsequently transforms downstream channel characteristics. These structures act as sediment traps, creating a &#x201c;sediment-starved&#x201d; condition downstream where rivers carry substantially reduced sediment loads. This cleaner water possesses greater erosive energy and enhanced capacity to scour riverbeds and banks. Additionally, dams reduce the geomorphic complexity of river systems by diminishing active channel areas and floodplain extent, while also modifying channel sinuosity and affecting the development and dimensions of sediment deposits. Conversely, upstream areas typically experience channel widening and sediment accumulation as the dominant geomorphological processes.</p>
<fig id="F7" position="float">
<label>FIGURE 7</label>
<caption>
<p>Conceptual model comparing the channel adjuctments in upstream and downstream of dam.</p>
</caption>
<graphic xlink:href="fenvs-13-1626165-g007.tif">
<alt-text content-type="machine-generated">Diagram illustrating morphological effects of a dam reservoir on a river. Upstream shows widening and aggradation with unexplored ecological consequences, referenced by twenty percent of literature. Downstream shows incision and narrowing with widely documented ecological consequences, referenced by eighty percent of literature.</alt-text>
</graphic>
</fig>
<p>Our analysis of morphological changes in upstream and downstream reaches of dammed rivers highlights significant differences in both the degree of understanding and the nature of channel adjustments. Morphological changes upstream of dams are significantly less documented than downstream one. The discrepancy in the number of studies between upstream and downstream dam sections can be attributed to the greater spatial extent of downstream dam effects, which range from a few to hundreds of kilometers, compared to the smaller upstream impact, typically from a few to dozens of kilometers (cf. e.g., <xref ref-type="bibr" rid="B58">Williams et al., 1984</xref>; <xref ref-type="bibr" rid="B31">Liro et al., 2020</xref>). The scientific focus on downstream adjustments largely reflects their pronounced and often negative impacts on river hydromorphology and aquatic ecosystems, as widely documented (<xref ref-type="bibr" rid="B45">Petts, 1979</xref>; <xref ref-type="bibr" rid="B46">1980a</xref>; <xref ref-type="bibr" rid="B21">Knighton, 1998</xref>; <xref ref-type="bibr" rid="B48">Petts and Gurnell, 2005</xref>; <xref ref-type="bibr" rid="B10">Gordon and Meentemeyer, 2006</xref>; <xref ref-type="bibr" rid="B54">Takahashi and Nakamura, 2011</xref>; <xref ref-type="bibr" rid="B13">Grant, 2012</xref>; <xref ref-type="bibr" rid="B36">Ma et al., 2012</xref>; <xref ref-type="bibr" rid="B52">Smith et al., 2016</xref>; <xref ref-type="bibr" rid="B51">Smith and Mohrig, 2017</xref>; <xref ref-type="bibr" rid="B5">Chong et al., 2021</xref>). Kondolf (1997) attributes these effects to sediment-starved flows &#x2014; often described as &#x201c;hungry water&#x201d; &#x2014; which cause severe bed and bank erosion.</p>
<p>Nevertheless, our review reveals a growing interest in upstream effects in recent years (e.g., <xref ref-type="bibr" rid="B63">Xu and Shi, 1997</xref>; <xref ref-type="bibr" rid="B38">Maselli et al., 2016</xref>; <xref ref-type="bibr" rid="B35">Luo et al., 2018</xref>; <xref ref-type="bibr" rid="B55">Volke et al., 2019</xref>; <xref ref-type="bibr" rid="B18">Hosseiny and Smith, 2019</xref>; <xref ref-type="bibr" rid="B31">Liro et al., 2020</xref>; <xref ref-type="bibr" rid="B15">Hamidifar and Nones, 2023</xref>). Some studies suggest that these adjustments may have positive effects on specific river hydromorphological characteristics (<xref ref-type="bibr" rid="B29">Liro, 2016</xref>; <xref ref-type="bibr" rid="B57">Wiejaczka and Kijowska-Struga&#x142;a, 2015</xref>). It is known that upstream dam adjustments occur over a smaller spatial area than downstream effects. However, some observed changes (e.g., the formation of channel bars in previously incised channels) may compensate for the loss of natural channel forms downstream, especially in regulated rivers (e.g., <xref ref-type="bibr" rid="B53">Surian and Rinaldi, 2003</xref>; <xref ref-type="bibr" rid="B60">Wy&#x17c;ga, 2007</xref>; <xref ref-type="bibr" rid="B24">Korpak, 2007</xref>; <xref ref-type="bibr" rid="B34">Lu et al., 2007</xref>; <xref ref-type="bibr" rid="B20">Khaleghi and Surian, 2019</xref>). Still, the ecological implications and long-term significance of upstream changes remain poorly understood, especially when compared to downstream effects (<xref ref-type="bibr" rid="B30">Liro, 2019</xref>; <xref ref-type="bibr" rid="B31">Liro et al., 2020</xref>). Addressing this knowledge gap is essential for developing comprehensive river management and dam impact mitigation strategies.</p>
<p>The literature review reveals that dam impact assessments have employed diverse methodological approaches and analytical tools. However, as noted by <xref ref-type="bibr" rid="B5">Chong et al. (2021)</xref>, we have entered an era characterized by extensive data collection capabilities and semi-automated geomorphological surveying techniques that offer new opportunities for examining morphological channel transformations. Consistent with Chong et al.&#x27;s findings, our review found no studies utilizing high-resolution data sources (such as drone imagery) or implementing repeated topographic surveys through systematic approaches like the Geomorphic Change Detection (GCD) methodology to quantify morphological alterations following dam construction.</p>
<p>Dam-induced channel adjustments have environmental and ecosystem effects, some of the possible positive consequence in upstream are including increased channel-floodplain connectivity, formation of valuable habitats in river channel (bars, channel island) (Wu et al., 2018), decrease of groundwater deficit and potential for protecting rejuvenated habitats and biota (<xref ref-type="bibr" rid="B33">Lobera et al., 2017</xref>; Liro et al., 2022), in downstream are potential for controlled increase of the river baseflow during water deficit periods, potential for partly reducing flood peaks (<xref ref-type="bibr" rid="B22">Kondolf and Swanson, 1993</xref>). Whereas, the negative effects in upstream are intensive fine sediments deposition on fish habitats, negative effects of prolonged artificial flooding biota and riparian vegetation (Sanchis-Ibor1 et al., 2018), potential for contaminated sediment accumulation, increased flood risk, a need to manage accelerated accumulation of contaminated sediments, a limited potentential for landuse due to artificial flooding (Liro et al., 2022). In terms of downstream, the possible negative effects are decrease in channel-floodplain connectivity, disappearance of habitats connected to sediment presence in river channel (bars, channel island), increase of groundwater defict, negative effects of hydropeaking on river fauna, intensivied bnak erosion resulting from channel incision (<xref ref-type="bibr" rid="B49">Phillips et al., 2005</xref>; <xref ref-type="bibr" rid="B19">Kale and Ataol, 2021</xref>), increase flood risk resulting from changes in channel geometry. Considering the issues raised and in terms of changes in river morphology caused by the dam, despite some positive ecological and management effects especially upstream of the dam, the negative effects of the dam are numerous, so building the dam is not recommended.</p>
<p>Finally, this work represents the first review about morphological changes of the river due to dam in compare upstream and downstream, worldwide. The study and data turned out to be neither homogeneous nor complete for all the study rivers that affect by dam. Some major limitations of this work are: (i) few data about bed-level changes, especially about channel aggradation, (ii) few case studies in some region that have large river and large dams, and, (iii) not equal and homogeneity in number of studies in different climatic regions. Consequently, providing definitive assessments of how river morphology responds to dam construction across different climatic regions remains challenging. Such comparative analyses are further complicated by variations in dam design specifications and operational parameters, as well as scale-related issues when comparing rivers of different magnitudes. While distinguishing between natural processes and anthropogenic influences (such as dams) on river morphological changes is typically difficult, our understanding of dam-induced channel modifications could be substantially enhanced through additional research efforts.</p>
</sec>
<sec sec-type="conclusion" id="s5">
<title>5 Conclusion</title>
<p>Despite the widespread global human intervention through dam construction, there remains a significant gap in the literature concerning river morphological changes and channel adjustments induced by dams. The majority of existing studies have predominantly focused on the impacts of dams and reservoirs on flow regimes and sediment transport, rather than on channel morphology. Our systematic review highlights distinct patterns of morphological adjustments occurring upstream and downstream of dams. Most studies have concentrated on downstream sections of dam, with considerably fewer studies addressing upstream morphological changes. Nonetheless, the primary downstream responses are channel incision and narrowing, whereas upstream adjustments are characterized mainly by channel widening and aggradation. These findings provide critical insights for assessing the future geomorphic and ecological consequences of dam construction and offer valuable guidance for the management of alluvial rivers influenced by dams. To enhance understanding and promote sustainable river management, future research should broaden their spatial scope beyond Asia and downstream reaches, incorporate emerging remote sensing and analytical technologies, and investigate the ecological implications of upstream morphological changes more comprehensively. It is worth noting that in studying and investigating the effects of dams on channel morphology in future research, two points should be considered. First, the effects of dams should be seen simultaneously with other factors affecting the channel, such as drainage basin conditions (e.g., climate, geology, and land use), and second, the effects of these morphological changes on ecology and management should be examined.</p>
</sec>
</body>
<back>
<sec sec-type="data-availability" id="s6">
<title>Data availability statement</title>
<p>The original contributions presented in the study are included in the article/<xref ref-type="sec" rid="s12">Supplementary Material</xref>, further inquiries can be directed to the corresponding author.</p>
</sec>
<sec sec-type="author-contributions" id="s7">
<title>Author contributions</title>
<p>SK: Supervision, Formal Analysis, Writing &#x2013; original draft, Writing &#x2013; review and editing, Software, Data curation, Investigation, Methodology, Project administration, Visualization, Resources, Conceptualization, Validation. KN: Conceptualization, Writing &#x2013; review and editing, Methodology.</p>
</sec>
<sec sec-type="funding-information" id="s8">
<title>Funding</title>
<p>The author(s) declare that no financial support was received for the research and/or publication of this article.</p>
</sec>
<ack>
<p>The authors gratefully acknowledge Dr. Esmaeel Saeedy Robat for his valuable assistance and advice during the systematic review process. We also thank Dr. Suiji Wang for kindly providing cross-sectional data of the lower Yellow River reach.</p>
</ack>
<sec sec-type="COI-statement" id="s9">
<title>Conflict of interest</title>
<p>The authors declare that the research was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest.</p>
</sec>
<sec sec-type="ai-statement" id="s10">
<title>Generative AI statement</title>
<p>The author(s) declare that no Generative AI was used in the creation of this manuscript.</p>
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<title>Publisher&#x2019;s note</title>
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<sec sec-type="supplementary-material" id="s12">
<title>Supplementary material</title>
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</p>
<supplementary-material xlink:href="Supplementaryfile1.pdf" id="SM1" mimetype="application/pdf" xmlns:xlink="http://www.w3.org/1999/xlink"/>
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