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<article xml:lang="EN" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" article-type="research-article">
<front>
<journal-meta>
<journal-id journal-id-type="publisher-id">Front. Clim.</journal-id>
<journal-title>Frontiers in Climate</journal-title>
<abbrev-journal-title abbrev-type="pubmed">Front. Clim.</abbrev-journal-title>
<issn pub-type="epub">2624-9553</issn>
<publisher>
<publisher-name>Frontiers Media S.A.</publisher-name>
</publisher>
</journal-meta>
<article-meta>
<article-id pub-id-type="doi">10.3389/fclim.2021.789869</article-id>
<article-categories>
<subj-group subj-group-type="heading">
<subject>Climate</subject>
<subj-group>
<subject>Original Research</subject>
</subj-group>
</subj-group>
</article-categories>
<title-group>
<article-title>ENSO Amplitude Asymmetry in Met Office Hadley Centre Climate Models</article-title>
</title-group>
<contrib-group>
<contrib contrib-type="author" corresp="yes">
<name><surname>Ineson</surname> <given-names>Sarah</given-names></name>
<xref ref-type="aff" rid="aff1"><sup>1</sup></xref>
<xref ref-type="corresp" rid="c001"><sup>&#x0002A;</sup></xref>
<uri xlink:href="http://loop.frontiersin.org/people/1505780/overview"/>
</contrib>
<contrib contrib-type="author">
<name><surname>Dunstone</surname> <given-names>Nick J.</given-names></name>
<xref ref-type="aff" rid="aff1"><sup>1</sup></xref>
</contrib>
<contrib contrib-type="author">
<name><surname>Ren</surname> <given-names>Hong-Li</given-names></name>
<xref ref-type="aff" rid="aff2"><sup>2</sup></xref>
<uri xlink:href="http://loop.frontiersin.org/people/1333561/overview"/>
</contrib>
<contrib contrib-type="author">
<name><surname>Renshaw</surname> <given-names>Richard</given-names></name>
<xref ref-type="aff" rid="aff1"><sup>1</sup></xref>
</contrib>
<contrib contrib-type="author">
<name><surname>Roberts</surname> <given-names>Malcolm J.</given-names></name>
<xref ref-type="aff" rid="aff1"><sup>1</sup></xref>
</contrib>
<contrib contrib-type="author">
<name><surname>Scaife</surname> <given-names>Adam A.</given-names></name>
<xref ref-type="aff" rid="aff1"><sup>1</sup></xref>
<xref ref-type="aff" rid="aff3"><sup>3</sup></xref>
</contrib>
<contrib contrib-type="author">
<name><surname>Yamazaki</surname> <given-names>Kuniko</given-names></name>
<xref ref-type="aff" rid="aff1"><sup>1</sup></xref>
</contrib>
</contrib-group>
<aff id="aff1"><sup>1</sup><institution>Met Office Hadley Centre, Met Office</institution>, <addr-line>Exeter</addr-line>, <country>United Kingdom</country></aff>
<aff id="aff2"><sup>2</sup><institution>State Key Laboratory of Severe Weather, Institute of Tibetan Plateau and Polar Meteorology, Chinese Academy of Meteorological Sciences</institution>, <addr-line>Beijing</addr-line>, <country>China</country></aff>
<aff id="aff3"><sup>3</sup><institution>College of Engineering, Mathematics and Physical Sciences, University of Exeter</institution>, <addr-line>Exeter</addr-line>, <country>United Kingdom</country></aff>
<author-notes>
<fn fn-type="edited-by"><p>Edited by: Swadhin Kumar Behera, Japan Agency for Marine-Earth Science and Technology (JAMSTEC), Japan</p></fn>
<fn fn-type="edited-by"><p>Reviewed by: Han-Ching Chen, University of Hawaii at Manoa, United States; Tomomichi Ogata, Japan Agency for Marine-Earth Science and Technology (JAMSTEC), Japan; Agus Santoso, University of New South Wales, Australia</p></fn>
<corresp id="c001">&#x0002A;Correspondence: Sarah Ineson <email>sarah.ineson&#x00040;metoffice.gov.uk</email></corresp>
<fn fn-type="other" id="fn001"><p>This article was submitted to Predictions and Projections, a section of the journal Frontiers in Climate</p></fn></author-notes>
<pub-date pub-type="epub">
<day>20</day>
<month>12</month>
<year>2021</year>
</pub-date>
<pub-date pub-type="collection">
<year>2021</year>
</pub-date>
<volume>3</volume>
<elocation-id>789869</elocation-id>
<history>
<date date-type="received">
<day>05</day>
<month>10</month>
<year>2021</year>
</date>
<date date-type="accepted">
<day>29</day>
<month>11</month>
<year>2021</year>
</date>
</history>
<permissions>
<copyright-statement>Copyright &#x000A9; 2021 Ineson, Dunstone, Ren, Renshaw, Roberts, Scaife and Yamazaki.</copyright-statement>
<copyright-year>2021</copyright-year>
<copyright-holder>Ineson, Dunstone, Ren, Renshaw, Roberts, Scaife and Yamazaki</copyright-holder>
<license xlink:href="http://creativecommons.org/licenses/by/4.0/"><p>This is an open-access article distributed under the terms of the Creative Commons Attribution License (CC BY). The use, distribution or reproduction in other forums is permitted, provided the original author(s) and the copyright owner(s) are credited and that the original publication in this journal is cited, in accordance with accepted academic practice. No use, distribution or reproduction is permitted which does not comply with these terms.</p></license>
</permissions>
<abstract><p>Long climate simulations with the Met Office Hadley Centre General Circulation Model show weak El Ni&#x000F1;o-Southern Oscillation (ENSO) amplitude asymmetry between El Ni&#x000F1;o and La Ni&#x000F1;a phases compared with observations. This lack of asymmetry is explored through the framework of a perturbed parameter experiment. Two key hypotheses for the lack of asymmetry are tested. First, the possibility that westerly wind burst activity is biased is explored. It is found that the observed difference in wind burst activity during El Ni&#x000F1;o and La Ni&#x000F1;a tends to be underestimated by the model. Secondly, the warming due to subsurface non-linear advection is examined. While the model exhibits non-linear dynamic warming during both La Ni&#x000F1;a and El Ni&#x000F1;o, and thus a contribution to ENSO asymmetry, it is shown to be consistently underestimated in comparison with ocean reanalyses. The non-linear zonal advection term contributes most to the deficiency and the simulation of the anomalous zonal currents may be playing a key role in its underestimation. Compared with the ocean reanalyses, the anomalous zonal currents associated with ENSO are too weak in the vicinity of the equatorial undercurrent and the surface wind driven zonal currents extend too deep.</p></abstract>
<kwd-group>
<kwd>El Ni&#x000F1;o-Southern Oscillation</kwd>
<kwd>asymmetry</kwd>
<kwd>non-linear dynamic warming</kwd>
<kwd>westerly wind burst</kwd>
<kwd>climate model</kwd>
</kwd-group>
<counts>
<fig-count count="5"/>
<table-count count="0"/>
<equation-count count="3"/>
<ref-count count="55"/>
<page-count count="10"/>
<word-count count="6607"/>
</counts>
</article-meta>
</front>
<body>
<sec sec-type="intro" id="s1">
<title>Introduction</title>
<p>El Ni&#x000F1;o-Southern Oscillation (ENSO) is the strongest natural interannual climate signal in the tropics, with oscillations between warm El Ni&#x000F1;o and cold La Ni&#x000F1;a phases occurring every few years. ENSO events show considerable spatio-temporal complexity (Timmermann et al., <xref ref-type="bibr" rid="B49">2018</xref>). In particular, the distribution of sea surface temperature (SST) variability in the Pacific is not Gaussian, in that the strongest El Ni&#x000F1;o events are stronger than the strongest La Ni&#x000F1;a events in the east Pacific (e.g., Burgers and Stephenson, <xref ref-type="bibr" rid="B6">1999</xref>). This amplitude asymmetry in SSTs is associated with asymmetry of temperatures in the ocean subsurface (Jin et al., <xref ref-type="bibr" rid="B23">2003</xref>; Zhang et al., <xref ref-type="bibr" rid="B55">2009</xref>; Hayashi and Jin, <xref ref-type="bibr" rid="B17">2017</xref>). There is also asymmetry in the duration of El Ni&#x000F1;o and La Ni&#x000F1;a events (Okumura and Deser, <xref ref-type="bibr" rid="B36">2010</xref>), but here we focus on the amplitude asymmetry as it is an outstanding model error (Zhang and Sun, <xref ref-type="bibr" rid="B54">2014</xref>).</p>
<p>ENSO amplitude asymmetry is generally underestimated in CMIP models (Sun et al., <xref ref-type="bibr" rid="B48">2016</xref>; Hayashi et al., <xref ref-type="bibr" rid="B18">2020</xref>; McKenna et al., <xref ref-type="bibr" rid="B33">2020</xref>). Because the largest El Ni&#x000F1;o events are often associated with severe impacts (Hardiman et al., <xref ref-type="bibr" rid="B16">2018</xref>; Anderson et al., <xref ref-type="bibr" rid="B4">2019</xref>), this deficiency impacts on our ability to make long range predictions of the risk of extreme regional events. It has also been suggested that future mean SST change in the east tropical Pacific may depend on a change in the asymmetry between the magnitudes of El Ni&#x000F1;o and La Ni&#x000F1;a SST anomalies under climate change (Ham, <xref ref-type="bibr" rid="B14">2017</xref>; Kohyama and Hartmann, <xref ref-type="bibr" rid="B27">2017</xref>). Better simulation of asymmetry may therefore also potentially improve future climate projections of ENSO (Hayashi et al., <xref ref-type="bibr" rid="B18">2020</xref>; Cai et al., <xref ref-type="bibr" rid="B7">2021</xref>).</p>
<p>While the reasons for ENSO amplitude asymmetry remain uncertain, a number of hypotheses have been put forward. For example, the westward shift of zonal wind stress anomalies during La Ni&#x000F1;a relative to El Ni&#x000F1;o may lead to relatively weak SST anomalies during La Ni&#x000F1;a (Kang and Kug, <xref ref-type="bibr" rid="B24">2002</xref>; Im et al., <xref ref-type="bibr" rid="B22">2015</xref>). A number of studies have also suggested an important role for westerly wind burst (WWB) events (Kessler and Kleeman, <xref ref-type="bibr" rid="B25">2000</xref>; Eisenman et al., <xref ref-type="bibr" rid="B11">2005</xref>; Chen et al., <xref ref-type="bibr" rid="B8">2015</xref>; Levine et al., <xref ref-type="bibr" rid="B28">2016</xref>; Hayashi and Watanabe, <xref ref-type="bibr" rid="B19">2017</xref>). Ocean non-linear dynamic warming either in the mixed layer (Jin et al., <xref ref-type="bibr" rid="B23">2003</xref>; An, <xref ref-type="bibr" rid="B3">2009</xref>; Su et al., <xref ref-type="bibr" rid="B47">2010</xref>) or ocean subsurface (Hayashi et al., <xref ref-type="bibr" rid="B18">2020</xref>; Chen et al., <xref ref-type="bibr" rid="B9">2021</xref>) is also thought to contribute. Similarly, another source of asymmetric heat convergence into the cold tongue region is associated with ocean tropical instability waves which occur preferentially in La Ni&#x000F1;a phases (Vialard et al., <xref ref-type="bibr" rid="B50">2001</xref>; An, <xref ref-type="bibr" rid="B2">2008</xref>). There is also asymmetry in the shortwave surface heat flux feedback (Lloyd et al., <xref ref-type="bibr" rid="B30">2012</xref>; Bayr et al., <xref ref-type="bibr" rid="B5">2018</xref>). For a recent review of these hypotheses see An et al. (<xref ref-type="bibr" rid="B1">2020</xref>), and references therein.</p>
<p>Here, we use the framework of a perturbed parameter experiment (PPE) of climate simulations (Yamazaki et al., <xref ref-type="bibr" rid="B53">2021</xref>) to explore the reasons for the weak asymmetry simulated in the Met Office Hadley Centre model, HadGEM-GC3.1 (Williams et al., <xref ref-type="bibr" rid="B52">2018</xref>). ENSO amplitude and periodicity have been shown to be sensitive to, for example, the specification of convection entrainment rate (Watanabe et al., <xref ref-type="bibr" rid="B51">2011</xref>; Lu and Ren, <xref ref-type="bibr" rid="B31">2016</xref>), and ENSO stability can be sensitive to the mean state (Kim et al., <xref ref-type="bibr" rid="B26">2014</xref>). The parameter combinations in each member of the PPE used in this study have been shown to yield a range of plausible yet diverse climate model behaviors (Sexton et al., <xref ref-type="bibr" rid="B46">2021</xref>; Yamazaki et al., <xref ref-type="bibr" rid="B53">2021</xref>). This paper focuses on two of the key hypotheses relating to ENSO asymmetry; the role of westerly wind bursts, and the model representation of subsurface ocean warming due to non-linear advection.</p>
</sec>
<sec sec-type="methods" id="s2">
<title>Methods</title>
<sec>
<title>Skewness</title>
<p>The ENSO amplitude asymmetry is typically measured by skewness, which is a measure of the extent to which a distribution differs from a normal distribution:</p>
<disp-formula id="E1"><label>(1)</label><mml:math id="M1"><mml:mrow><mml:mtext>Skewness&#x000A0;</mml:mtext><mml:mo>=&#x000A0;</mml:mo><mml:mfrac><mml:mrow><mml:msub><mml:mtext>m</mml:mtext><mml:mn>3</mml:mn></mml:msub></mml:mrow><mml:mrow><mml:msup><mml:mrow><mml:mo stretchy='false'>(</mml:mo><mml:msub><mml:mtext>m</mml:mtext><mml:mn>2</mml:mn></mml:msub><mml:mo stretchy='false'>)</mml:mo></mml:mrow><mml:mrow><mml:mn>3</mml:mn><mml:mo>/</mml:mo><mml:mn>2</mml:mn></mml:mrow></mml:msup></mml:mrow></mml:mfrac></mml:mrow></mml:math></disp-formula>
<p>where <italic>m</italic><sub><italic>k</italic></sub> is the <italic>k</italic>th moment,</p>
<disp-formula id="E2"><mml:math id="M2"><mml:mrow><mml:msub><mml:mtext>m</mml:mtext><mml:mtext>k</mml:mtext></mml:msub><mml:mo>&#x000A0;</mml:mo><mml:mo>=</mml:mo><mml:mo>&#x000A0;</mml:mo><mml:mstyle displaystyle='true'><mml:munderover><mml:mo>&#x02211;</mml:mo><mml:mrow><mml:mtext>i</mml:mtext><mml:mo>=</mml:mo><mml:mtext>1</mml:mtext></mml:mrow><mml:mtext>N</mml:mtext></mml:munderover><mml:mrow><mml:mfrac><mml:mrow><mml:msup><mml:mrow><mml:mo stretchy='false'>(</mml:mo><mml:msub><mml:mtext>x</mml:mtext><mml:mtext>i</mml:mtext></mml:msub><mml:mtext>&#x000A0;</mml:mtext><mml:mo>&#x02212;</mml:mo><mml:mtext>&#x000A0;</mml:mtext><mml:mover accent='true'><mml:mtext>X</mml:mtext><mml:mo stretchy='true'>&#x000AF;</mml:mo></mml:mover><mml:mo stretchy='false'>)</mml:mo></mml:mrow><mml:mtext>k</mml:mtext></mml:msup></mml:mrow><mml:mtext>N</mml:mtext></mml:mfrac></mml:mrow></mml:mstyle></mml:mrow></mml:math></disp-formula>
<p>where <italic>x</italic><sub><italic>i</italic></sub> is the ith observation, the overbar denotes the mean, and <italic>N</italic> the number of observations.</p>
<p>Here, skewness is calculated from monthly data.</p>
</sec>
<sec>
<title>Non-linear Advection</title>
<p>The ocean heat budget can be expressed as:</p>
<disp-formula id="E3"><label>(2)</label><mml:math id="M3"><mml:mtable columnalign='left'><mml:mtr><mml:mtd><mml:mfrac><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:msup><mml:mi>T</mml:mi><mml:mo>&#x02032;</mml:mo></mml:msup></mml:mrow><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:mi>t</mml:mi></mml:mrow></mml:mfrac><mml:mtext>&#x02009;</mml:mtext><mml:mo>=</mml:mo><mml:mtext>&#x02009;</mml:mtext><mml:mo>&#x02212;</mml:mo><mml:mrow><mml:mo>(</mml:mo><mml:mrow><mml:msup><mml:mi>u</mml:mi><mml:mo>&#x02032;</mml:mo></mml:msup><mml:mfrac><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:mover accent='true'><mml:mi>T</mml:mi><mml:mo>&#x000AF;</mml:mo></mml:mover></mml:mrow><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:mi>x</mml:mi></mml:mrow></mml:mfrac><mml:mtext>&#x02009;</mml:mtext><mml:mo>+</mml:mo><mml:mtext>&#x02009;</mml:mtext><mml:msup><mml:mi>v</mml:mi><mml:mo>&#x02032;</mml:mo></mml:msup><mml:mfrac><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:mover accent='true'><mml:mi>T</mml:mi><mml:mo>&#x000AF;</mml:mo></mml:mover><mml:mi>x</mml:mi></mml:mrow><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:mi>y</mml:mi></mml:mrow></mml:mfrac><mml:mo>+</mml:mo><mml:msup><mml:mi>w</mml:mi><mml:mo>&#x02032;</mml:mo></mml:msup><mml:mfrac><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:mover accent='true'><mml:mi>T</mml:mi><mml:mo>&#x000AF;</mml:mo></mml:mover></mml:mrow><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:mi>z</mml:mi></mml:mrow></mml:mfrac><mml:mtext>&#x02009;</mml:mtext><mml:mo>+</mml:mo><mml:mtext>&#x02009;</mml:mtext><mml:mover accent='true'><mml:mi>u</mml:mi><mml:mo>&#x000AF;</mml:mo></mml:mover><mml:mfrac><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:msup><mml:mi>T</mml:mi><mml:mo>&#x02032;</mml:mo></mml:msup></mml:mrow><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:mi>x</mml:mi></mml:mrow></mml:mfrac><mml:mtext>&#x02009;</mml:mtext><mml:mo>+</mml:mo><mml:mtext>&#x02009;</mml:mtext><mml:mover accent='true'><mml:mi>v</mml:mi><mml:mo>&#x000AF;</mml:mo></mml:mover><mml:mfrac><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:msup><mml:mi>T</mml:mi><mml:mo>&#x02032;</mml:mo></mml:msup></mml:mrow><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:mi>y</mml:mi></mml:mrow></mml:mfrac></mml:mrow></mml:mrow></mml:mtd></mml:mtr><mml:mtr><mml:mtd><mml:mtext>&#x02009;&#x02009;&#x02009;&#x02009;&#x02009;&#x02009;&#x02009;&#x02009;&#x02009;&#x02009;&#x02009;&#x02009;&#x02009;&#x02009;&#x02009;</mml:mtext><mml:mrow><mml:mrow><mml:mo>+</mml:mo><mml:mtext>&#x02009;</mml:mtext><mml:mover accent='true'><mml:mi>w</mml:mi><mml:mo>&#x000AF;</mml:mo></mml:mover><mml:mfrac><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:msup><mml:mi>T</mml:mi><mml:mo>&#x02032;</mml:mo></mml:msup></mml:mrow><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:mi>z</mml:mi></mml:mrow></mml:mfrac></mml:mrow><mml:mo>)</mml:mo></mml:mrow><mml:mo>&#x02212;</mml:mo><mml:mrow><mml:mo>(</mml:mo><mml:mrow><mml:msup><mml:mi>u</mml:mi><mml:mo>&#x02032;</mml:mo></mml:msup><mml:mfrac><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:msup><mml:mi>T</mml:mi><mml:mo>&#x02032;</mml:mo></mml:msup></mml:mrow><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:mi>x</mml:mi></mml:mrow></mml:mfrac><mml:mtext>&#x02009;&#x02009;</mml:mtext><mml:mo>+</mml:mo><mml:msup><mml:mi>v</mml:mi><mml:mo>&#x02032;</mml:mo></mml:msup><mml:mfrac><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:msup><mml:mi>T</mml:mi><mml:mo>&#x02032;</mml:mo></mml:msup></mml:mrow><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:mi>y</mml:mi></mml:mrow></mml:mfrac><mml:mo>+</mml:mo><mml:msup><mml:mi>w</mml:mi><mml:mo>&#x02032;</mml:mo></mml:msup><mml:mfrac><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:msup><mml:mi>T</mml:mi><mml:mo>&#x02032;</mml:mo></mml:msup></mml:mrow><mml:mrow><mml:mo>&#x02202;</mml:mo><mml:mi>z</mml:mi></mml:mrow></mml:mfrac></mml:mrow><mml:mo>)</mml:mo></mml:mrow><mml:mo>+</mml:mo><mml:msup><mml:mi>R</mml:mi><mml:mo>&#x02032;</mml:mo></mml:msup></mml:mtd></mml:mtr></mml:mtable></mml:math></disp-formula>
<p>Where <italic>T</italic>, <italic>u</italic>, <italic>v</italic>, and <italic>w</italic> are temperature, zonal, meridional, and vertical velocities, respectively. The overbar and prime denote climatological mean and anomalies, respectively. The first bracket represents the linear terms, the second bracket is the nonlinear terms and the residual term, R, denotes thermodynamic and subgrid-scale contributions. The heat budget terms are estimated from monthly mean anomalies with respect to the long-term monthly climatology.</p>
</sec>
<sec>
<title>Models</title>
<p>ENSO asymmetry is assessed in a number of long climate simulations of the Met Office Hadley Centre general circulation model HadGEM-GC3.1 (Williams et al., <xref ref-type="bibr" rid="B52">2018</xref>) in three different resolutions: low (LL), medium (MM), and high (HH). LL and MM are CMIP6 preindustrial control simulations, at N96 (atmosphere midlatitude spacing of 135 km)/ORCA1 (1&#x000B0; ocean with enhanced meridional resolution near the equator) and N216 (60 km)/ORCA025 (1/4&#x000B0;) resolution, respectively (Menary et al., <xref ref-type="bibr" rid="B35">2018</xref>). Each run length is 500 years. HH has a resolution of N512 (25 km)/ORCA12 (1/12&#x000B0;) and forcing representative of the year 1950 (Roberts et al., <xref ref-type="bibr" rid="B43">2019</xref>). The run length is 150 years. All configurations have 85 levels in the atmosphere and 75 levels in the ocean. To investigate possible reasons for the weak asymmetry in these models, we use a PPE based on a similar model, HadGEM-GC3.05, at MM (N216/ORCA025) resolution. The objective of a PPE is to represent model uncertainty by perturbing model parameters within realistic ranges. There are 19 members, the &#x0201C;standard variant&#x0201D; (STD) unperturbed member (member 0) and 18 members with combinations of perturbations applied to atmosphere and land surface parameters. No perturbations are applied to the ocean. We use 200-year simulations with CMIP5 greenhouse gas and radiative forcing for the year 1900. Heat and fresh-water flux adjustments are derived for STD and applied to all PPE members. In addition, for the 18 perturbed members, a globally uniform, member specific, heat flux adjustment is applied (Yamazaki et al., <xref ref-type="bibr" rid="B53">2021</xref>). The methodology behind the choice of the perturbed parameters and the values they take is described in detail by Sexton et al. (<xref ref-type="bibr" rid="B46">2021</xref>), and the identifier which links the ensemble members to the parameter perturbations is shown in brackets: 0 (00000), 1 (00090), 2 (00605), 3 (00834), 4 (01113), 5 (01554), 6 (01649), 7 (01843), 8 (01935), 9 (02089), 10 (02123), 11 (02242), 12 (02305), 13 (02335), 14 (02491), 15 (02753), 16 (02832), 17 (02868), 18 (02914).</p>
</sec>
<sec>
<title>WWB Index</title>
<p>WWBs are identified here as events where the daily 10 m zonal wind anomaly, averaged from 2.5&#x000B0;N-2.5&#x000B0;S, exceeds 3 ms<sup>&#x02212;1</sup> over a longitude range of at least 10&#x000B0; and lasts for at least 3 consecutive days in the equatorial Pacific (130&#x000B0;E-80&#x000B0;W). We define a WWB index as the sum of the maximum anomaly for each event over a given time period to give a measure of accumulated strength. In recent literature, a number of different criteria have been used for defining WWBs, leading to varying results in terms of numbers of WWBs identified (Santoso et al., <xref ref-type="bibr" rid="B44">2017</xref>). The criteria used here gives an observed average WWB count of just over 12 per year which is within the range identified by other studies (e.g., Puy et al., <xref ref-type="bibr" rid="B40">2016</xref>).</p>
</sec>
<sec>
<title>Verification Data</title>
<p>Model data are compared with monthly mean SST from HadISST1.1 for 1873 to 2020 (Rayner et al., <xref ref-type="bibr" rid="B42">2003</xref>), monthly zonal wind stress from ERA5 for 1950 to 2020 (Hersbach et al., <xref ref-type="bibr" rid="B21">2020</xref>), and monthly mean depth of the 20&#x000B0; isotherm estimated from the ocean temperature dataset, EN4, for 1950 to 2020 (Good et al., <xref ref-type="bibr" rid="B12">2013</xref>). WWB activity is calculated using daily 10 m winds from the ERA-Interim reanalysis for 1980 to 2017 (Dee et al., <xref ref-type="bibr" rid="B10">2011</xref>). Monthly mean ocean potential temperature and currents are from two products, the ocean reanalysis used in the GloSea5 seasonal forecast system (MacLachlan et al., <xref ref-type="bibr" rid="B32">2015</xref>) and a recent ocean reanalysis based on the GO5 ocean model (Megann et al., <xref ref-type="bibr" rid="B34">2014</xref>), both for the period 1993 to 2017. In the scatter diagrams, the time-period used for calculating HadISST skewness is chosen to be the same as that used for the corresponding reanalysis variable.</p>
</sec>
</sec>
<sec sec-type="results" id="s3">
<title>Results</title>
<sec>
<title>Skewness in Hadley Centre Models</title>
<p>Assessment of ENSO characteristics for HadGEM-GC3.1 at MM resolution shows that this model performs favorably with observations across a range of performance metrics (Williams et al., <xref ref-type="bibr" rid="B52">2018</xref>) and in historical simulations, the PPE shows a range of plausible ENSO variability (Yamazaki et al., <xref ref-type="bibr" rid="B53">2021</xref>). In this study, standard deviation for Nino3.4 (170-120&#x000B0;W, 5&#x000B0;N-5&#x000B0;S) across the PPE ranges from 0.51&#x000B0;C to 0.97&#x000B0;C. El Ni&#x000F1;o and La Ni&#x000F1;a composite SST anomaly patterns in HadGEM-GC3.1 are also reasonably realistic and similar at MM and HH resolutions, although for LL the ENSO pattern extends somewhat too far west (Menary et al., <xref ref-type="bibr" rid="B35">2018</xref>; Roberts et al., <xref ref-type="bibr" rid="B43">2019</xref>). However, compared with the observations, these El Ni&#x000F1;o and La Ni&#x000F1;a patterns are too symmetrical, reflecting a lack of extreme El Ni&#x000F1;o events in the models.</p>
<p>Amplitude asymmetry, measured here by SST skewness is weaker than in observations in all our model simulations and versions, irrespective of resolution, as shown in <xref ref-type="fig" rid="F1">Figure 1</xref>. Because skewness in the more commonly used Ni&#x000F1;o3 region (150-90&#x000B0;W, 5&#x000B0;N-5&#x000B0;S) is weak in the PPE, we focus on a region in the far east Pacific (FEP) (110-80&#x000B0;W, 5&#x000B0;N-5&#x000B0;S) that maximizes skewness in the observations (Su et al., <xref ref-type="bibr" rid="B47">2010</xref>; Santoso et al., <xref ref-type="bibr" rid="B44">2017</xref>).</p>
<fig id="F1" position="float">
<label>Figure 1</label>
<caption><p>Comparison of skewness in Met Office Hadley Centre models. Skewness in 3 PPE members <bold>(A&#x02013;C)</bold> and observations (HadISST, 1873-2020) <bold>(D)</bold>. Rectangular box shows skewness in the far eastern Pacific region (FEP, 110-80&#x000B0;W, 5&#x000B0;N-5&#x000B0;S). Skewness in FEP for PPE members (blue), high, medium, and low resolution HadGEM-CG3.1 simulations (brown), and observations (black) <bold>(E)</bold>. Large circles show skewness over the full period of each model simulation, vertical bars show the 16th and 84th percentiles (corresponding to 1 standard deviation from the mean) and small circles are the maximum and minimum of skewness for 50-year sliding windows.</p></caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fclim-03-789869-g0001.tif"/>
</fig>
<p>The spatial pattern of SST skewness for the most positively skewed (SK&#x0002B;), standard variant (STD), and least positively skewed (SK&#x02013;) members of the 19-member PPE (<xref ref-type="fig" rid="F1">Figures 1A&#x02013;C</xref>) can be compared with the observations (<xref ref-type="fig" rid="F1">Figure 1D</xref>). STD shows a similar pattern of skewness but is much weaker than observed, with an FEP skewness of 0.34 compared with 1.02 for observations. SK- shows weakly negative skewness along the equator with an FEP skewness of &#x02212;0.11. For SK&#x0002B;, our most skewed simulation, the westward extent of the region of positive skewness is similar to that observed but again the SK&#x0002B; member skewness for the FEP region is much weaker than observed, 0.65 compared with 1.02, and the pattern is more equatorially confined.</p>
<p>Skewness for the FEP region for all PPE members shows diversity across the ensemble in the mean, and also a variety of multidecadal variability (<xref ref-type="fig" rid="F1">Figure 1E</xref>). However, no member has a mean skewness as high as observed, or indeed a maximum skewness in any 50-year period as high as the observed mean. We also show model data from different resolution HadGEM-GC3.1 models with lower (LL), the same (MM), and higher (HH) ocean and atmosphere resolution as the PPE. These all show similar characteristics to STD and lie within the PPE range, suggesting that lack of ENSO asymmetry is insensitive to resolution in HadGEM-GC3.1</p>
<p>Observations are most strongly positively skewed in the far east Pacific, decreasing to become weakly negatively skewed in the region near the dateline and in a horseshoe-like pattern in the west Pacific and extending somewhat into the South Pacific Convergence zone and the Intertropical Convergence zone. We note that PPE member SK&#x0002B; has unrealistically high negative skewness in these regions and that in the PPE there is a tendency for the more positively skewed members to also have unrealistically high negative skewness in the west Pacific.</p>
</sec>
<sec>
<title>Mean State</title>
<p>In the tropical Pacific there is a close relationship between mean equatorial SST, zonal wind stress and thermocline depth in the mean state (Philander, <xref ref-type="bibr" rid="B38">1990</xref>). As expected, the PPE reflects this, with significant correlations between these variables across individual member simulations.</p>
<p>The relationship between these key variables and skewness also shows significant correlations, with higher SST (<italic>r</italic> = 0.54, <italic>p</italic> &#x0003C; 0.02), weaker zonal wind stress (<italic>r</italic> = 0.57, <italic>p</italic> &#x0003C; 0.02) and shallower 20&#x000B0;C isotherm in the west pacific (<italic>r</italic> = &#x02212;0.44, <italic>p</italic> &#x0003C; 0.1) all being related to higher skewness in the model (<xref ref-type="fig" rid="F2">Figure 2</xref>). The PPE members with deeper thermocline in the western Pacific tend to have a somewhat shallower thermocline in the eastern Pacific (<xref ref-type="fig" rid="F2">Figure 2E</xref>). Biases in the coupled model mean state can therefore clearly impact the model skewness and in principle could be responsible for the lack of ENSO asymmetry. However, as the observations lie well within the range of the modeled mean state for each of these assessed variables, the model mean state is not systematically biased in this respect. A realistic representation of these variables by the model does not in itself therefore necessarily lead to realistic asymmetry and we reject the hypothesis that these mean state errors are responsible for the lack of ENSO asymmetry.</p>
<fig id="F2" position="float">
<label>Figure 2</label>
<caption><p>Relationship between skewness and equatorial Pacific annual mean state. Spatial plot of grid point correlation of SST <bold>(A)</bold> zonal wind stress <bold>(C)</bold> and depth of 20&#x000B0; isotherm <bold>(E)</bold> with FEP skewness. Scatter diagram of FEP skewness against annual mean Ni&#x000F1;o3 SST <bold>(B)</bold>, central Pacific wind stress <bold>(D)</bold> and depth of 20&#x000B0; isotherm for region Ni&#x000F1;o4 <bold>(F)</bold> for PPE members (small blue circles, with SK&#x0002B; magenta, STD red, SK- cyan) and reanalysis data (large black circle). Regions for FEP (110-80&#x000B0;W, 5&#x000B0;N-5&#x000B0;S), Ni&#x000F1;o3 (150-90&#x000B0;W, 5&#x000B0;N-5&#x000B0;S), central Pacific (160&#x000B0;E-80&#x000B0;W, 5&#x000B0;N-5&#x000B0;S) and Ni&#x000F1;o4 (160&#x000B0;E-150&#x000B0;W, 5N-5S) are shown by boxes in the left-hand panel.</p></caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fclim-03-789869-g0002.tif"/>
</fig>
</sec>
<sec>
<title>Westerly Wind Events</title>
<p>The relationship between WWBs and skewness is examined using a simple index as a measure of accumulated wind burst activity (WWB index, see section Methods) The index is calculated from daily 10 m zonal winds that have previously been filtered to remove timescales longer than a season (90 days).</p>
<p>We find there are large differences in the annual mean WWB index across our model versions, ranging from an average of 18.3 to 108.8 ms<sup>&#x02212;1</sup> per year (<xref ref-type="fig" rid="F3">Figure 3A</xref>). However, there is only a weak (not significant) positive correlation (<italic>r</italic> = 0.28) between FEP skewness and WWB activity. We also note that although the two members with largest skewness have relatively high levels of WWB activity, we see that this is not a prerequisite for the PPE skewness. Indeed, the member with the lowest WWB index has similar moderate skewness to the member with the highest WWB index. Finally, we note that the WWB index for the reanalysis lies well within the range of the PPE simulations, suggesting that the model versions are not systematically biased with respect to annual mean WWB activity.</p>
<fig id="F3" position="float">
<label>Figure 3</label>
<caption><p>Relationship between skewness and westerly wind burst activity. Scatter diagram of FEP SST skewness against annual mean wind burst activity <bold>(A)</bold> and against the El Ni&#x000F1;o minus La Ni&#x000F1;a difference in wind burst activity during the January to November period prior to ENSO events <bold>(B)</bold> for PPE members (small blue circles, with SK&#x0002B; magenta, STD red, SK&#x02013; cyan) and reanalysis data (large black circle). Horizontal bars show the standard error.</p></caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fclim-03-789869-g0003.tif"/>
</fig>
<p>To test the importance of the modulation of wind burst activity by SST in the PPE we look at the difference in WWB activity between El Ni&#x000F1;o and La Ni&#x000F1;a events in the January to November period prior to ENSO events (<xref ref-type="fig" rid="F3">Figure 3B</xref>). El Ni&#x000F1;o/La Ni&#x000F1;a events are defined here as DJF Ni&#x000F1;o3.4 anomalies exceeding a threshold of &#x0002B;/- 0.8&#x000B0;C. As has been previously demonstrated (e.g., Puy et al., <xref ref-type="bibr" rid="B40">2016</xref>) there is a significant modulation of WWBs by ENSO, with more WWBs during El Ni&#x000F1;o. However, the majority of PPE members show only a weak difference, significantly underestimating the observed modulation. We also look at this relationship for two separate periods, during boreal spring, early in the ENSO growth cycle, and in boreal summer-autumn, when SST anomalies are established and growing in magnitude. We find a similar relationship for both periods (not shown), suggesting that the rather weak difference in WWB activity in the PPE may result in very large initial El Ni&#x000F1;o growth in the PPE being less likely than observed, and also, in the second part of the year, that the model may be underestimating the observed dependence of WWBs on SST. However, because of the lack of extreme El Ni&#x000F1;o events in the PPE this latter result may also simply reflect this lack of model skewness. While there is no relationship (<italic>r</italic> = 0.2) across the PPE with skewness, the difference between the model and observations suggests possible model error in HadGEM-GC3.1 in this respect.</p>
</sec>
<sec>
<title>Warming Due to Ocean Non-linear Advection</title>
<p>Equatorial non-linear advection for the combined ocean reanalysis product shows warming in the region of the thermocline and in the surface mixed layer, largely concentrated to the east of the dateline (<xref ref-type="fig" rid="F4">Figure 4A</xref>). These features agree well with the analysis shown by Hayashi et al. (<xref ref-type="bibr" rid="B18">2020</xref>). Non-linear advection for the PPE, combined over all models, shows a similar pattern, but with muted warming in the ocean subsurface (<xref ref-type="fig" rid="F4">Figure 4B</xref>).</p>
<fig id="F4" position="float">
<label>Figure 4</label>
<caption><p>Warming due to non-linear advection. Non-linear advection (K month<sup>&#x02212;1</sup>) for combined ocean analyses <bold>(A)</bold> and combined PPE members <bold>(B)</bold>. Box shows the averaging region for the non-linear advection. Scatter diagram of FEP skewness against non-linear advection for PPE ensemble members (small blue circles, with SK&#x0002B; magenta, STD red, SK&#x02013; cyan) and ocean reanalyses (large black circles) <bold>(C)</bold>. Non-linear advection terms during growth period of El Ni&#x000F1;o (red) and La Ni&#x000F1;a (blue) for combined ocean reanalyses <bold>(D)</bold> combined PPE <bold>(E)</bold>. Combined results are the average over the respective datasets.The growth period is defined here as the average of months April to November prior to the event, with the DJF El Ni&#x000F1;o/La Ni&#x000F1;a events identified using a &#x000B1;0.8&#x000B0;C threshold. Data in <bold>(C)</bold> are area-averages taken over the box shown in the upper panels (180&#x000B0;-100&#x000B0;W,1&#x000B0;S-1&#x000B0;N, 50&#x02013;150 m). The combined ocean analysis data <bold>(A)</bold> have been smoothed. Vertical bars in <bold>(D,E)</bold> show the 95% confidence interval.</p></caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fclim-03-789869-g0004.tif"/>
</fig>
<p>Following Hayashi et al. (<xref ref-type="bibr" rid="B18">2020</xref>), area averages are calculated for the region (180-100&#x000B0;W, 1&#x000B0;S-1&#x000B0;N, 50&#x02013;150 m), encompassing the maximum subsurface non-linear dynamic warming (NDW) in the reanalysis. There is a significant relationship (<italic>r</italic> = 0.74, <italic>p</italic> &#x0003C; 0.001) between the FEP skewness and the NDW for the model simulations and ocean reanalyses (<xref ref-type="fig" rid="F4">Figure 4C</xref>). The correlation over PPE members alone is also significant (<italic>r</italic> = 0.41, <italic>p</italic> &#x0003C; 0.1). Unlike our other hypothesized mechanisms for the underestimated ENSO asymmetry, there is a clear systematic model bias in the strength of the NDW and all of the PPE members underestimate the NDW relative to the ocean reanalysis. The NDW is therefore a strong candidate for the lack of asymmetry in the model, consistent with other studies on NDW (Hayashi et al., <xref ref-type="bibr" rid="B18">2020</xref>; Chen et al., <xref ref-type="bibr" rid="B9">2021</xref>).</p>
<p>To consider the contribution of NDW to the growth of ENSO, the NDW for the mean of April to November prior to the peak of ENSO is calculated for all El Ni&#x000F1;o and La Ni&#x000F1;a events. Overall, there is a positive contribution during <italic>both</italic> El Ni&#x000F1;o and La Ni&#x000F1;a for both the combined ocean reanalysis and the model members (<xref ref-type="fig" rid="F4">Figures 4D,E</xref>) This acts to weaken cold (La Ni&#x000F1;a) events and strengthen warm (El Ni&#x000F1;o) events, contributing to amplitude asymmetry. However, the total warming in the reanalysis is over twice the magnitude of that in the model (see the right-hand bars in <xref ref-type="fig" rid="F4">Figures 4D,E</xref>).</p>
<p>We now consider the three component terms of the NDW (Equation 2) to try to better understand the origin of the weak NDW in the model. The zonal non-linear warming term (-u&#x00027;&#x02202;T&#x00027;/&#x02202;x) and meridional term (-v&#x00027;&#x02202;T&#x00027;/&#x02202;y) both make a positive contribution to the warming, whereas the vertical (-w&#x00027;&#x02202;T&#x00027;/&#x02202;z) term is negative and acts to partially offset the horizontal terms. Comparing the PPE with the ocean analysis we see that it is the zonal component of non-linear advection which largely accounts for the difference in total magnitude (left hand bars in <xref ref-type="fig" rid="F4">Figures 4D,E</xref>). It is noteworthy that there is also a significant relationship (<italic>r</italic> = 0.48, <italic>p</italic> &#x0003C; 0.05) between skewness and NDW for the zonal component across PPE members (not shown).</p>
<p>To further explore the reason for the difference in zonal NDW, we decomposed the zonal NDW for the observational reanalysis and the model (<xref ref-type="fig" rid="F5">Figures 5A,B</xref>), into the zonal anomalous temperature gradient (<xref ref-type="fig" rid="F5">Figures 5C,D</xref>) and anomalous zonal current (<xref ref-type="fig" rid="F5">Figures 5E,F</xref>), although this separation is an approximation due to the correlation between the strength of T&#x00027; and u&#x00027; fluctuations. It can be seen that the anomalous eastward surface current anomalies extend too deep in the PPE compared with the reanalysis and in particular, that the anomalous westward currents in the region of the equatorial undercurrent are much weaker in the model (<xref ref-type="fig" rid="F5">Figures 5E,F</xref>). This latter difference suggests that the ocean circulation response to ENSO is too weak in our model and provides a focus for future model development.</p>
<fig id="F5" position="float">
<label>Figure 5</label>
<caption><p>Decomposition of zonal component of non-linear advection. Zonal component of non-linear advection (K month<sup>&#x02212;1</sup>) for combined ocean reanalysis <bold>(A)</bold> and combined PPE <bold>(B)</bold>, zonal gradient of temperature anomaly (K m<sup>&#x02212;1</sup>) for combined ocean reanalysis <bold>(C)</bold> and combined PPE <bold>(D)</bold>, and zonal current anomaly (ms<sup>&#x02212;1</sup>) for combined ocean reanalysis <bold>(E)</bold> and combined PPE <bold>(F)</bold>. Sections are for 1&#x000B0;N-1&#x000B0;S, for the growth period of ENSO, El Ni&#x000F1;o plus La Ni&#x000F1;a for <bold>(A,B)</bold> and El Ni&#x000F1;o minus La Ni&#x000F1;a for <bold>(C&#x02013;F)</bold>. The combined ocean analysis data have been smoothed.</p></caption>
<graphic mimetype="image" mime-subtype="tiff" xlink:href="fclim-03-789869-g0005.tif"/>
</fig>
</sec>
</sec>
<sec sec-type="discussion" id="s4">
<title>Discussion</title>
<p>We have tested several hypotheses for the weak ENSO amplitude asymmetry in HadGEM-GC3.1, finding initially that mean state errors do not appear to be a factor limiting skewness in our model, at least in the large-scale areas examined here.</p>
<p>Overall, there is only a weak relationship between skewness and mean WWB activity and our model is not systematically biased in this respect. There is also no significant relationship between the state dependent WWB activity and skewness which is consistent with the recent study by Olson et al. (<xref ref-type="bibr" rid="B37">2021</xref>) who used an empirical stochastic model to suggest that state dependent noise does not play a role in El Ni&#x000F1;o asymmetry. However, in this case most of the PPE ensemble members do underestimate the difference in WWB activity between El Ni&#x000F1;o and La Ni&#x000F1;a, suggesting the likelihood of model error in this respect. This is consistent with Levine et al. (<xref ref-type="bibr" rid="B28">2016</xref>) who show that most CMIP5 models underestimate state-dependent noise forcing. Nevertheless, our simple diagnostic ignores the spatio-temporal diversity of westerly wind bursts (Hao et al., <xref ref-type="bibr" rid="B15">2019</xref>) which could in principle affect the oceanic impact (Puy et al., <xref ref-type="bibr" rid="B40">2016</xref>) and the SST response to WWBs may also depend on the oceanic background state, and hence be affected by ocean model bias (Puy et al., <xref ref-type="bibr" rid="B41">2019</xref>) so a more sophisticated analysis would be worthwhile to confirm our results.</p>
<p>In contrast, variation in subsurface NDW is significantly related to skewness and it is also underestimated in our model compared with observational ocean reanalyses. Our analysis suggests model bias in the subsurface zonal NDW, and particularly in anomalous zonal ocean currents which are too weak. These findings are in agreement with other recent analyses of CMIP5 and CMIP6 models (Hayashi et al., <xref ref-type="bibr" rid="B18">2020</xref>; Chen et al., <xref ref-type="bibr" rid="B9">2021</xref>).</p>
<p>The reasons for the weak ENSO response in the equatorial undercurrent variability are not fully understood, especially as the mean undercurrent strength is reasonably represented in the 0.25&#x000B0; ocean model. It&#x00027;s possible there may be a number of contributing factors. First, the mean ENSO amplitude in the combined PPE is slightly lower (by about 10%) than in the ocean reanalysis period, which would suggest slightly weaker amplitude ocean variability in the PPE. Secondly, and in common with other climate models, the dynamic coupling, usually measured by the regression on Ni&#x000F1;o4 zonal stress onto Ni&#x000F1;o3 SST, is underestimated (Planton et al., <xref ref-type="bibr" rid="B39">2021</xref>). In the PPE the regression coefficient is from 15 to 55% lower than observed, indicating that the ENSO related momentum transfer to the ocean is too weak. In addition, the anomalous upper ocean currents extend somewhat too deep compared with the reanalysis, perhaps indicating that there is too much mixing of momentum in the upper ocean.</p>
<p>The mechanisms tested here do not rule out other explanations. However, at the 0.25&#x000B0; ocean model resolution used in our simulations, heat convergence associated with tropical instability waves is in reasonably good agreement with observational estimates (Graham, <xref ref-type="bibr" rid="B13">2014</xref>), suggesting that tropical instability waves are unlikely to be the factor limiting skewness in these models. In addition, we note that in our model there is no significant change in skewness across resolution. Other mechanisms have not been tested. We note that model members that have too cold mean SST in the east Pacific also tend to have lower skewness, which is consistent with Sun et al. (<xref ref-type="bibr" rid="B48">2016</xref>), who find that weak non-linear air&#x02013;sea interaction due to cold tongue bias limits asymmetry. Similarly, Liang et al. (<xref ref-type="bibr" rid="B29">2017</xref>), find that the range of physical parameters for a system to fall into a regime with positive asymmetry may be rather narrow.</p>
<p>The Met Office Hadley Centre uses the same model in seasonal, decadal and climate prediction systems. While on seasonal timescales the initialized prediction realistically captures the amplitude of extreme El Ni&#x000F1;o events (Scaife et al., <xref ref-type="bibr" rid="B45">2017</xref>), on longer timescales the predictions revert to the control behavior (Hermanson et al., <xref ref-type="bibr" rid="B20">2018</xref>) with no asymmetry and too few, if any, extreme El Ni&#x000F1;o events. This may impact our ability to predict and evaluate the risk of extreme regional events associated with the most intense El Ni&#x000F1;os and highlights the importance of model assessment (e.g., Planton et al., <xref ref-type="bibr" rid="B39">2021</xref>) and further analysis of this important model error, with a view to future improvement.</p>
</sec>
<sec sec-type="data-availability" id="s5">
<title>Data Availability Statement</title>
<p>The original contributions presented in the study are included in the article/supplementary material, further inquiries can be directed to the corresponding author.</p>
</sec>
<sec id="s6">
<title>Author Contributions</title>
<p>KY, MR, and RR gave advice on, the PPE, HH, and ocean reanalysis data, respectively. SI carried out the analysis. AS, ND, and H-LR contributed to discussing the results and writing the paper. All authors commented on the manuscript.</p>
</sec>
<sec sec-type="funding-information" id="s7">
<title>Funding</title>
<p>This work, and contributors SI, ND, and AS, were supported by the UK-China Research &#x00026; Innovation Partnership Fund through the Met Office Climate Science for Service Partnership (CSSP) China as part of the Newton Fund. H-LR was supported by China National Science Foundation (Grant No. 41975094) and the National Key Research and Development Program of China (Grant No.2018YFC1506000). KY was supported by Department for Business, Energy and Industrial Strategy. MR was supported by, and the HH model was performed as part of, the Horizon 2020 program PRIMAVERA project (Grant No. 641727). This study has been conducted using E.U. Copernicus Marine Service Information.</p>
</sec>
<sec sec-type="COI-statement" id="conf1">
<title>Conflict of Interest</title>
<p>The authors declare that the research was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest.</p>
</sec>
<sec sec-type="disclaimer" id="s8">
<title>Publisher&#x00027;s Note</title>
<p>All claims expressed in this article are solely those of the authors and do not necessarily represent those of their affiliated organizations, or those of the publisher, the editors and the reviewers. Any product that may be evaluated in this article, or claim that may be made by its manufacturer, is not guaranteed or endorsed by the publisher.</p>
</sec>
</body>
<back>
<ref-list>
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